286 
Psyche 
[December 
Moscow Branch of the All-Union Entomological Society, Academy 
of Sciences of the U. S. S. R., compared specimens of 5 Mammoth 
Cave trechines (all except P. audax ) with the 3 examples of An- 
ophthalmus striatus j 3 A. menetriesii, and 2 A. ventricosus in the 
Motschulskv collection. Dr. Arnoldi (in litt.) reports that the con- 
ventional interpretation of striatus and menetriesii , with ventricosus 
a synonym of menetriesii (Jeannel, 1928), is correct. 
P. striatus is a riparian species in Mammoth Cave, apparently feed- 
ing on tubificid annelids in the silt banks along Lake Lethe and Echo 
River. In August, 1965, T. G. Marsh and I collected 30 Pseudan- 
ophthalmus at the margin of Lake Lethe, in dim illumination beside 
the electrically lighted tourist trail. Only one specimen of this series 
was P. menetriesii and the remainder were P. striatus. On the other 
hand, menetriesii predominates in the upper levels of the cave, where 
most of the older collections seem to have been made. 
In a study of distribution and variation in the menetriesii group 
Barr and Marsh (in preparation) have found that the range of 
menetriesii is a narrow belt not far removed from the Dripping 
Spring escarpment. P. striatus , on the other hand, extends eastward 
across Barren County into the southwest corner of Metcalf County, 
where it is locally larger, more convex, and more difficult to separate 
morphologically from menetriesii. In Metcalf County striatus is 
sympatric with another, undescribed species of the menetriesii group. 
There is good evidence for character displacement where the range 
of striatus overlaps that of menetriesii or that of the undescribed 
species. 
P. audax was known only from the type series (Barr, 1959), col- 
lected about 1880 in a cave 9 miles east of Mammoth Cave, until 
it was taken in White Cave, a half mile south-southwest of the His- 
toric Entrance to Mammoth Cave, in August, 1961 (1 cf), and 
August, 1965 (1 $). The species occurred on wet, rotting wood at 
the back of White Cave, near an old bridge across a shallow pit. 
The pit is supposedly separated from Crevice Pit in Mammoth Cave 
only by a narrow crevice impenetrable to man. P. menetriesii , P. 
puhescens , P. inexpectatus , P. striatus, and N. tellkampfii have all 
been found at this place in the cave, either on the White Cave side 
of the crevice, the Mammoth Cave side, or both. Although I sug- 
gested that audax was probably “a rather curious offshoot from the 
menetriesii-robustus branch of the genus” (Barr, 1959: p. 3), further 
study of the genus has shown that this species should form the type 
of an audax group (as suggested by Jeannel, 1949), to include P. 
