1982] Holldobler & Engel- Siegel — Glands in Male Ants 123 
Fig. 3 Longitudinal section through pygidial gland of Pogonomyrmex barbatus 
S- GC=glandular cells; R=reservoir. 
Sternal glands: 
More than in any other subfamily, the males of the doryline ants 
are also richly endowed with sternal glands (Tab. 1; Fig. 4). We 
found major complex glands with intersegmental reservoirs between 
the 3rd and 4th, 4th and 5th, 5th and 6th sternites. Glandular clus- 
ters are also present between the 6th and 7th sternites — but without 
a pronounced reservoir. In addition, clusters of glandular cells 
whose ducts penetrate the sclerotized cuticle were found in the 3rd 
sternite, they are also strongly developed in the 8th and 9th sternites 
(Fig. 4). Similar sternal glands were found in the 8th sternite of 
males of Myrmecia pilosula, Pachycondvla apiaealis and P. obscuri- 
cornis. In both Pachycondvla species the glandular ducts open in 
bundles into cuticular cups located in the 8th sternite (Fig. 7a). 
In the ponerine species Leptogenys diminuta the males possess a 
huge intersegmental sternal gland between the 7th and 8th sternites. 
This gland consists of large paired clusters of glandualr cells. Each 
cell sends a duct into wider collecting channels which lead into a 
paired large reservoir, consisting of ventro-lateral invaginations of 
the intersegmental membrane (Fig. 6). There is a second paired 
sternal gland between the 8th and 9th sternite; but this gland is 
