DESCRIPTION OF A NEW SPECIES OF 
KROM BEINIUS (H YMENOPTERA: PERILAMPIDAE) 
FROM THE PHILIPPINES, AND THE PHYLOGENETIC 
RELATIONSHIPS OF THE GENUS* 
Bv D. Christopher Darling 
Department of Entomology, 
Cornell University, Ithaca, N.Y. 14853 
The genus Krombeinius (Hymenoptera: Perilampidae) was re- 
cently described (Boucek 1978) to include perilampids with an 
amalgam of the characters of Euperi/ampus Latreille and Peri/am- 
pus Walker. The habitus suggests Euperi/ampus, and there are two 
synapomorphies to unite these two genera (Darling 1983): postspi- 
racular sclerite reduced to a narrow triangle, less than one-half as 
wide as the adjacent pronotum; and pronotum massive, at least 
one-third the length of the mesoscutum. However, Krombeinius 
exhibits the wing venation, presence of a marginal rim on the scutel- 
lum, and large third metasomal tergite characteristic of Perilampus. 
I regard these as plesiomorphic similarities. The genus is character- 
ized by the absence of the defining apomorphic characteristics of 
Euperi/ampus, i.e., by symplesiomorphy. 
In this paper I present new information on the structure of the 
male genitalia and labrum of the type species of Krombeinius. These 
structures have proved to be of considerable value in defining gen- 
era in the Perilampidae (Darling 1983). From this analysis 1 suggest 
autapomorphies for defining Krombeinius. In addition, I describe a 
new species of Krombeinius from the Philippines, and discuss the 
affinities of the three included species. 
Taxonomic studies of Krombeinius have been hampered by the 
scarcity of material. The type species, K. eumenidarum, was de- 
scribed by Boucek (1978) from a series of specimens (2 male, 2 
female) reared from the larvae ol an eumenine wasp in Sri Lanka. 
All specimens were prematurely killed and had to be liberated from 
the pupal cuticles, producing some abnormalities in the type mate- 
rial. Also included by Boucek (1978) in this genus was Perilampus 
megalaspis Cameron, known only from the type material (3 females) 
* Manuscript received by the editor September 22, 1982. 
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