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[Vol. 87 
Fig. 2. A-B, Aimed discharges elicited by pinching a left midleg (A) and left 
hindleg (B) of Necrodes with forceps. The spray pattern is visible on phenophthalein 
indicator paper. C-F, Directional aiming movements of the anal turret of Necrodes. 
Note that the abdominal tip is accurately pointed toward the site of application of the 
stimulus: (C) tibia of midleg, (D) tarsus of midleg, (E) tibia of hindleg, pinched with 
forceps; (F) back of beetle touched with hot spatula. 
tests on phenolphthalein indicator paper and had the recognizable 
terpenoid fragrance of the spray. 
Examination of fresh spray ejected by Necrodes on glass showed 
occasional presence of opaque material in the discharged fluid, sug- 
gesting that the secretion may sometimes be expelled with admix- 
ture of fecal paste. Since the glandular contents are forced to the 
outside by way of the rectum, such admixture may occur whenever 
the pathway of secretory egress is blocked by enteric matter. Two 
fatty acids not listed in Table 1, stearic acid and palmitic acid, were 
