60 BULLETIN 57, UNITED STATES NATIONAL MUSEUM. 
DESMALOPEX, new genus. 
Type-species. — Pteropus leucopterus Temminck (as understood by 
Dobson, Catal. Chiropt. Brit. Mus., p. 32. 1878). 
Geographical distribution . — Philippine Islands. 
Number of forms . — The type is the only species known. 
Characters— Like Pteropus / but skull with broader rostrum and 
palate, orbits directed slightly more upward, and teeth showing sev- 
eral peculiarities that suggest Pteralopex. Upper incisors subequal, 
distinctly larger than in Pteropus , the cross section of the crown 
nearly one-third that of canine, the cingulum produced into a notice- 
able shelf posteriorly. Lower 4 incisors very unequal, the crotvn area 
of the outer nearly one-half that of canine, that of the inner scarcely 
more than one-lialf as great. Canines not peculiar, the upper with- 
out secondary cusps. Small upper premolar well developed, not 
deciduous, its diameter nearly half that of upper incisor, its crown 
flat. Small lower premolar relatively larger than in Pteropus , but 
smaller than in Pteralopex , its outer edge raised but not distinctly 
cuspidate. The remaining premolars, both above and below, agree 
with those of Pteropus , except that pm s shows no trace of cusp on 
inner side. Molars, both above and below, subquadrate in outline, 
the length of the crown never conspicuously greater than the width 
(m 3 and m 1 not elongated as in Pteropus) . In the maxillary teeth 
the cusps are like those of Pteropus except that they are higher rel- 
atively to length of crown. Lower molars peculiar in that the 
ridges of m x and m, are each divided into two low but distinct 
rounded cusps. The quadritubercular form resulting from this is 
very noticeable in m 15 less so in m 2 . Third lower molar as in 
Pteropus. 
Species examined. — Desmalopex leucopterus (Temminck). 
Remarks . — This genus is intermediate between Pteropus and Pte- 
ralopex, though nearer the former. In the broadened rostrum and 
slightly upturned orbits the skull distinctly suggests Pteralopex , 
while the same tendency is shown by the form and relative size of the 
incisors, the well-developed small upper premolar, the squarish out- 
line of the molars, and the extra cusps of m \ and m 2 . Externally 
there are no characters worthy of note. 
Genus PTERALOPEX Thomas. 
1888. Pteralopex Thomas, Ann. and Mag. Nat. Hist., 6th ser., I, p. 155, 
February, 1888. ( atrata .) 
1899. Pteralopex Matschie, Flederm. des Berliner Mus. fiir Naturk., p. 11. 
(Subgenus of Pteropus.) 
Type-species. — Pteralopex atrata Thomas. 
Geographic distribution . — Solomon Islands. 
Number of forms— Only the type species has been thus far dis- 
covered. 
