THE FAMILIES AND GENERA OF BATS. 
65 
developed. There is a rudimentary papilla-like tail, the calcar is 
present, as is the claw of the index finger, the wings are attached low 
on the sides, as in Epomophorus , and the fur is slightly woolly. 
Species examined. — Scotonycteris bedfordi Thomas. 
Remarks. — This genus has been placed near Cynopterus by Mats- 
chie, but it appears to be more closely related to Epomophorus. 
Though the skull more nearly resembles that of the former, this char- 
acter seems to indicate merely a less specialized stage than has been 
reached by E pomophorus. Similarly primitive are the slightly de- 
veloped lips. The number of teeth, the form of the mandible, and 
the broadly hairy back all resemble E pomophorus , while the struc- 
ture of the teeth could easily be derived from that of the related 
genus by a process nearly the opposite to that which has taken place 
in Hypsignathus. 
Genus EPOMOPHORUS Bennett. 
1836. Epomophorus Bennett, Proc. Zool. Soc., London (1835), p. 149, Feb- 
ruary 12, 1836. ( macroceplialus .) 
1866. Epomops Gray, Proc. Zool. Soc., London, p. 65 ( franqueti ). 
1878. Epomophorus Dobson, Catal. Chiropt. Brit. Mus., p. 4 (part). 
1899. Epomophorus Matschie, Flederm. des Berliner Mus. fiir Naturk., p. 34 
(part). 
1899. Epomops Matschie, Flederm. des Berliner Mus. fiir Naturk., p. 56 
(subgenus for E. comptus, E. franqueti, and E. clohsonii). 
1899. Micropteropus Matschie, Flederm. des Berliner Mus. fiir Naturk., p. 57 
(subgenus for E. pusillus). 
1899. N anonycteris Matschie, Flederm. des Berliner Mus. fiir Naturk., p. 58 
(subgenus for E. veldkampii) . 
Type-speeies. — Pteropus epomophorus Bennett— /h macroceplialus 
Ogilby. 
Geographic distribution . — Africa, south of the Sahara. 
Number of forms . — About two dozen forms of Epomophorus are 
now known. Next to Pteropus this is the largest genus of Mega- 
chiroptera. 
Characters. — General appearance much as in Pteropus , but lips 
fuller; tail present, though usually not visible externally; occipital 
portion of skull (fig. 9) slightly deflected and not tubular, and upper 
cheek teeth only 3-3. Dental formula \ 
- 2 3. 1. -- 3 4 5 -- .2 - 2 1 - 1 2- 2 1 - 1 - 
12 1. -23156 - *2 - 2’ c l- V pm Z - 3 ’ m 2 - 2 -28, 
Upper incisors small, sty li form ; the crowns pointed, but not much 
differentiated from shafts; the outer tooth of each pair shorter than 
inner, but not strongly contrasted in size. The incisors are sepa- 
rated from each other by narrow spaces and from the canine by a 
wide diastema. Lower incisors equal in size, forming a nearly 
straight row between the canines, separated from each other and 
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