332 The Philippine Journal of Science i9i7 
here recognizes four species which in the order of their vegeta- 
tive development, the most amply developed first, are : 
1. C. Eberhardtii Christ. 
2. C. tricuspis (Hooker) Christ. 
3. C. Sagitta (Christ) Copel. 
4. C. cantoniensis (Baker) Christ. 
The sterile frond of C. Eberhardtii is large and five lobed. 
The fertile frond is unknown. In this paper, Christ suggests 
the relationship of Christiopteris and Cheiropleuria. 
Bower * has recently made a very careful anatomical study 
of the fern hitherto best known as Leptochilus tricuspis (Hooker) 
C. Chr. but which he calls “Gymnopteris {Leptochilus) tricuspis 
(Hook.), C. Chr.,” a name which may surprise Christensen. 
From his summary, I quote: 
3. L. tricuspis stands alone in the latter genus (Leptochilus) in various 
features, but especially in the diplodesmic character. It should, therefore, 
be removed, and by reviving its old generic name, now merged in Leptochilus, 
it may be styled Gymnopteris tricuspis (Hook.), Bedd. Of that genus it 
will be at present the only species. 
The name which Bower attempts to revive could not possibly 
stand as valid; but there is still no necessity for a new name, 
as that already cited, given by Christ and listed in the Supple- 
ment to Christensen’s Index, is certainly correct. 
As noted by Bower, Mrs. Schumann ” has grouped together 
Leptochilus tricuspis and L. varians (Mett.) Fournier, a fern 
known only in New Caledonia, noting that both have bilateral 
spores. The herbarium of the Bureau of Science has a number 
of specimens of the latter species, which struck me several years 
ago as by no possibility belonging in Leptochilus, and as probable 
relatives of Christiopteris. Bower’s paper has stimulated me to 
investigate this more carefully. There is a special bundle system 
underlying the sporangia, but this is not at all sharply distin- 
guished, in the fronds studied, from the general bundle system 
nearer the upper surface of the frond, which is homologous to 
the bundle system of the sterile frond. There are many small 
paraphyses which are always or nearly always simple. The 
paleae of the rhizome are ovate, with many or several border 
cells with thin walls, without trichome-like outgrowths, and not 
at all reddish in color. 
A similar examination of Christiopteris Sagitta has shown 
‘ Studies in the Phylogeny of the Filicales, VI. Ferns showing the 
‘Acrostochoid’ condition, with special reference to dipteroid derivatives, 
Ann. Bot. 21 (1917) 1-39. 
Die Acrosticheen und ihre Stellung im System der Fame, Flora 1 08 
(1915) 201-260. 
