xn,c, 6 Copeland: The Geyius Christiopteris 335 
or sagittate, caudate, with the marginal walls alone thin and 
with small teeth, which, hov/ever, have thick middle walls, being 
formed by the joint projection of two marginal cells and the 
wall between them. 
Cyclophorus angmtatus Desv., a species notable for its elon- 
gated sori, has a single bundle layer under the sorus. The spores 
are bilateral and the annulus is of 14 or 15 cells. The paleae 
are long and slender, but not caudate, entire and without very 
thin walls, although colorless except near the base. The hypo- 
dermis is of very large cells which are not very irregular. 
Photinopteris speciosa J. Sm. has an obvious and irregular 
hypodermis not essentially different from that characteristic of 
the Phymatodes group. Its spores are bilateral and the annulus 
is of about 13 cells. The leaf is much compressed under the 
sorus, without room for more than one bundle in a layer. The 
paleae are peltate, but immediately narrowed to a long, harsh, 
hair-like point. The marginal wall is usually thin, with very 
peculiar teeth. The smaller of these are like those of Drymo- 
glossum and Hymenolepis ; but there are present also others in 
which the teeth elongate, one of the cells sometimes outgrowing 
the other; or both cells may elongate, and separate in the outer 
part. Photinopteris has enough characters in common with 
Christiopteris so that their general affinity is unmistakable; 
although, as Bower has already shown, Christ’s suggestion that 
they may have eventually to be united is untenable. On the 
other hand, Photinopteris is likewise unmistakably a relative 
of the Phymatodes group; this is the only construction which 
I can place on the presence in both of the peculiar layer of cells 
under the upper epidermis. 
Aglaomorpho. pilosa Copel. (Dryostachyum pilosum J. Sm.) has 
reniform spores and setose sporangia with an annulus of 
14 or 15 cells. The lamina under the hymenium is very thin, 
without room for a diplodesmic vascular supply. The paleae 
are peltate at the base, and lanceolate, with the walls gradually 
becoming thinner toward the margin. The margin itself is 
laeerate-ciliate by separation from the body of the scale of the 
upper ends of the marginal cells. The hypodermis, already 
figured in my paper on comparative ecology,’ has the Phyma- 
todes characteristics. 
The affinity of Photinopteris speciosa J. Sm. and Aglaomorpha 
pilosa Copel. is unmistakable. The most peculiar characteristic 
of Photinopteris is probably the isolated gland below the base 
of each pinna. Aglaomorpha pilosa has likewise a glandular 
’ Philip. Journ. Sci. 2 (1907) Bot. 3, fig. 36. 
