44 
FOSSIL REPTILIA OF THE 
its articulation with the tympanic (28)- The metacarpus and dislocated nnguiculate digits 
of the wing-limb are confusedly interblended with the crushed and dislocated back part 
of this skull; three phalanges {ir \, ir^, jv 5) of the wing-finger are determinable. 
The two anterior teeth (!'. 2') of the mandible show longitudinal angular depressions at 
their base, indicating exposure of their inner side, and that they belong to the left ramus. 
The corresponding part of the right ramus may have been broken away ; the third laniary 
(3') clearly belongs to this ramus, which is fractured beneath its socket. The point of this 
tooth is broken off: Avhat remains of the body is curved, and is implanted more obliquely 
backward than the two preceding teeth. This at first led me to suspect it might be the 
foremost tootli of the mandible, and that the left ramus had been pushed in advance as well 
as downward : but my doubts on this point have been set at rest by the specimen (PL 
XVIII) next to be described, and I view the tooth in question as the third of the mandibular 
series : it is divided from the second by an interval of 6 lines, and the second stands at a 
rather shorter interval behind the first. Five lines behind the third tooth is the base of a 
fourth laniary (4'), and four lines further back is an indication of a fifth (.5'). This is followed 
by the characteristic series of between thirty and forty very small, subcompressed cuspidate 
teeth, each less than a line in length, corresponding in extent with the maxillary part of 
the upper jaw. The entire series of mandibular teeth occupies an extent of alveolar 
border measuring 5 inches 1 line. 
The depth of the right ramus gradually increases from 5 lines below the last laniary 
to 10 lines below the last denticle. The inner side of the dislocated ramus (32') shows a 
strong longitudinal ridge projecting inwards about 3 lines above the lower border. The 
outer surface of the ramus seems to have been strengthened near its lower border by a 
similar but lower ridge. 
The distal ends of the antibrachial bones (54, 55) overlap the hind part of the mandible : 
that which shows the larger articular surface, opposite the three slender metacarpals, 
should be the radius. The base of the supplementary styloid bone appears near the distal 
end of the ulna, but is better shown in Buckland’s original specimen.^ Indications of 
two carpals intervene between these and the metacarpus. This overlies and conceals the 
articular pedicle of the mandible and contiguous parts (squamosal, malar, &c.) of the 
skull. The metacarpus includes the three slender supports of digits 4 r/, rrq and the 
strong and thick metacarpal of the wing-finger (uA This bone, being almost con- 
cealed by the first phalanx in Buckland’s specimen, was overlooked, and that phalanx 
was described as the metacarpal of the wing-finger, which, accordingly, in the restoration, 
fig. 2, PI. 27, of ‘Buckland’s Memoir,’ is made three times the length of the other and 
more slender metacarpals (3'). I have, therefore,- had that part of the original specimen, 
now in the British Museum, redrawn (PI. XIX, fig. 1), the true metacarpal being shown at 
m 4. It corresponds with the same bone in previously described Pterosauria by surpassing 
in thickness, not in length, the other constituents of the metacarpus. In the specimen, 
1 As in the part re-drawn in fig. 1, PI. XIX. 
