jecting above eye in profile; 2 ors, 2 ori (some- 
times 3); gena narrow, 0.125 height of eye; epi- 
stoma lacking; 3+1 strong dc ; frons matt, sooty 
black; ocellar triangle and orbit weakly shining, 
lunule silvery; antennae black; mesonotum and 
scutellum blackish, weakly shining; squama gray, 
margin and fringe deep black; last and penultimate 
sections of vein M 3+4 variable, but last slightly 
longer or at most 1.5 times as long; male geni- 
talia with aedeagus as in figures 479, 480, 
distiphallus strongly pigmented; sperm pump with 
large blade and strongly developed base. 
Host/Early Stages. Unknown, possibly Salix. 
Distribution. California. 
Reference. Spencer, 1981: 134. 
14 (13). Small species, wing length 2. 5-2.6 mm 
P. calyptrata (Hendel) 
Synopsis. Agreeing closely with P. californic a but 
mesonotum more grayish; male genitalia with ae- 
deagus (figs. 481, 482) having distiphallus weak- 
ly pigmented. 
Host/Early Stages. Unknown, possibly Salix. 
Distribution. Illinois, Maryland; Canada. 
Reference. Part 2, p. 270. 
Large species, wing length in female 3.75 mm 
P . sp. ( Salix ) 
Synopsis. Agreeing closely with P. setosa but all 
antennal segments deep black, frons also uniformly 
black; male genitalia unconfirmed, possibly 
as in figure 483. 
Host/Early Stages. Almost certainly Salix spp. 
Distribution. New York, certainly more widespread. 
Reference. Part 2, p. 272. 
15 (12). Normally 4 orbital bristles, 2 ors reclinate 
- JP. setosa (Loew) 
Synopsis. Frons matt or at most slightly pro- 
jecting above eye in profile, sooty black but 
normally at least faintly reddish adjoining 
lunule, this large, silvery or frequently suffused 
with reddish; 3d antennal segment reddish brown, 
never entirely black; orbital bristles strong, 
normally 2 ors and 2 ori, occasionally a 3d ori on 
one side, more rarely on both; mouth margin with 
only narrow epistoma; mesonotum ash gray, with 3+1 
strong dc, prsc strong; legs largely black, knees 
only slightly paler; male genitalia with aedeagus 
having long paired distal tubules, only slightly 
diverging, virtually straight in side 
view (figs. 484, 485, holotype). 
Host/Early Stages. Acer rub rum , probably other 
Acer spp. Larva mining in trunk and root, leaving 
tree to pupate at ground level; larva full grown 
in July and August (Virginia), overwintering as 
puparium, adults emerging in April or May; length 
of larva 15-17 mm; puparium pale yellow, cylindri- 
cal, posterior spiracles each with 3 bulbs (fig. 
486, A, B). 
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