80 
- Male genitalia with aedeagus having distiphallus 
unpigmented (figs. 1241, 1244) 
79 (78). Mesopleuron only narrowly yellow along upper 
margin P. angel icae Kaltenbach 
Synopsis. Frons 2-2.5 times width of eye, 
bright yellow, hindmargin of eye black; upper ors 
always more slender than lower, normally 2 ori; 
gena about 0.33 height of eye; 3d antennal seg- 
ment small, round, black, 1st and 2d segments 
only slightly paler; mesopleuron dark, apart 
from narrow yellow upper margin; mesonotum matt 
grayish black, acr in 4 rows; squama yellowish, 
margin and fringe dark; wing length from 2 mm 
in male to 3 mm in female, 2d costal section 
normally about 3 times length of 4th; male 
genitalia with aedeagus (fig. 1235) having 
distiphallus with fully pigmented, 
slender paired tubules in same plane as basal 
sclerites, these without spicules (fig. 1236). 
Host/Early Stages. Most commonly Angelica spp., 
also Heracleum 1 ana turn ; in Europe Laserpitium , 
more rarely Aegopodium . Larva forming primary 
upper surface blotch mine (fig. 1237), several 
mines in same leaf may give appearance of 
single large mine; puparium black, posterior 
spiracles each with ellipse of about 20 minute 
bulbs . 
Distribution. California; Alaska, Europe. 
References. Griffiths, 1973a: 233; Spencer, 
1981: 373. 
- Mesopleuron yellow on upper quarter 
JP. astotinensis Griffiths 
Synopsis. Closely resembling angel icae but 
mesopleuron more broadly yellow, somewhat 
smaller, wing length up to 2.5 mm in female, 2d 
costal section shorter, about 2.5 times length 
of 4th; male genitalia with aedeagus having 
distiphallus only partially pigmented, lying 
approx, at right angles to basal sclerites, 
these with conspicuous spicules (figs. 1238, 
1239). 
Host/Early Stages. Solidago f lexicaulis (in 
Alberta, S_. canadensis , S_. gigantea ) . Larva 
forming irregular upper surface linear mine 
(fig. 1240); puparium black, posterior spir- 
acles each with 15-21 bulbs. 
Distribution. Wisconsin (new record); Canada. 
References. Griffiths, 1976b: 248; part 2, 
p. 310. 
80 (78). Male genitalia with aedeagus having small, chit- 
inized notch at lower 3d of distiphallus; small 
spiniform sclerite at end of right basal scle- 
rite (fig. 1241) JP. spondylii Robineau-Desvoidy 
Synopsis. Frons and gena orange yellow, with 
upper orbit slightly darkened, upper ors great- 
ly reduced (rarely lacking) ; 3d antennal seg- 
201 
