and tarsi largely dark; mesonotum blackish 
gray, only weakly shining, acr in 2 rows; 
squama gray, margin and fringe black; wing with 
2d costal section about 3 times length of 4th; 
male genitalia with aedeagus (fig. 1336) having 
2 rows of spinules toward distal end of basal 
sclerites; distiphallus directed dorsally, 
broadly fused at base (fig. 1337); sperm pump 
with large blade. 
Host/Early Stages. Sanicula crassicaulis (syn., 
S^. menziesii ) and S.. laciniata in California, S. 
marilandica in Alberta. Larva forming broad, 
irregular upper surface mine (fig. 1338), 
greenish when fresh, brown when older; puparium 
black, posterior spiracles each with ellipse of 
about 15 bulbs. 
Distribution. California; Canada. 
References. Spencer, 1969a: fig. 536; 1981: 
417. 
Note. Available material suggests that P. sani- 
culae is smaller than P. lanat i and JP. oenan - 
thoides , but positive identification of caught 
specimens may only be possible from the male 
genitalia. 
Larger species, wing length 2. 1-2.8 mm 129 
129 (128). 1 ori P. oenanthoides Spencer 
Synopsis. Closely resembling P. saniculae but 
somewhat larger, wing length 2. 1-2.4 mm, 2d 
costal section longer, 3-4 times length of 4th; 
acr more numerous, irregularly in 4 rows; male 
genitalia with aedeagus (fig. 1339) having 
spinules above both left and right basal scle- 
rites, those on left fewer but stronger, 
distiphallus largely unpigmented. 
Host/Early Stages. Oenanthe sarmentosa . Larva 
forming irregular upper surface linear mine 
(fig. 1340); puparium black, posterior spir- 
acles greatly enlarged (for aquatic environ- 
ment), 2-armed, with 15 bulbs on one arm, 10 on 
other (see I>. cicutae Hendel in Europe, Spencer, 
1976a: fig. 704). 
Distribution. California (Alameda Co.). 
Reference. Spencer, 1981: 406. 
2 ori 130 
130 (129). Male genitalia with aedeagus (figs. 1341, 1342) 
having strip of spinulate sclerotization beside 
ventral lobe P. lanati Spencer 
Synopsis. Generally resembling both oenan- 
thoides and P. saniculae , possibly larger, wing 
length 2. 1-2.8 mm; frons variable, either al- 
most completely yellow dorsally (see couplet 75) 
or more uniformly brown. 
Host/Early Stages. Primarily Osmorhiza , in- 
cluding 0. chilensis (syn., 0. nuda) , 0. 
longistylis , and 0. occidentalis , possibly 
Heracleum lanatum . Larva forming greenish 
219 
