linear mine, largely following serrations along 
leaf margin; puparium black. 
Distribution. California; Canada. 
References. Spencer, 1966b: 108; 1969a: 250; 
1981: 391. 
Note. This species was described when the host 
was believed to be Heracleum lanatum ; the nor- 
mal host was found to be Osmorhiza during 
further collecting in California in 1977. 
Male genitalia with aedeagus (fig. 1343) without 
secondary band of sclerotizat ion beside ventral 
lobe, this with both sides conspicuously ex- 
panded at ventral end osmorhizae Spencer 
Synopsis. Agreeing with P. lanati in external 
characters but frons apparently more consist- 
ently dark brown. 
Host/Early Stages. Osmorhiza claytonii and 0. 
longistylis . Larva forming upper surface 
linear mine (fig. 1344); puparium frequently 
lightly adhering to underside of leaf until 
dislodged by wind or rain. 
Distribution. Virginia; new record: Wisconsin, 
Dane Co., Madison, 1 m. ex Osmorhiza claytonii , 
27.V.75 (S. Tavormina). 
Reference. Spencer, 1969a: 261. 
131 (127). Frons brownish black or upper half paler, ochre- 
ous 132 
Frons uniformly black 135 
132 (131). Legs with femora entirely black 133 
At least foreknee yellow 134 
133 (132). Frons dark, uniformly brownish black 
j>. S onorensis Spencer 
Synopsis. Frons dark brown, orbit black; upper 
ors weak, slender, 2 ori; all antennal segments 
black, 3d slightly longer than broad; mesono- 
tum black, predominantly matt with weak sub- 
shine; wing length in male 2.1 mm, 2d costal 
section just over twice length of 4th; male 
genitalia with aedeagus as in figures 1345, 
1346; sperm pump large. 
Host/Early Stages. Unknown. 
Distribution. California (mountains). 
Reference. Spencer, 1981: 421. 
Frons paler above, ochreous P. columbinae Sehgal 
Synopsis. Frons black in lower half, orbit and 
lunule black; 2 ori, lower weak; antenna 
black, 3d segment small, round; mesonotum black, 
only weakly shining; wing length 1.5-1. 8 mm, 2d 
costal section twice length of 4th; male geni- 
talia with aedeagus as in figures 1347, 1348; 
sperm pump with narrow stalk and rounded blade. 
Host/Early Stages. Aquilegia spp., Thai ict rum 
polycarpum , and other Thalictrum spp. Larva 
forming blotch mine; puparium dark brown, 
covered with small spicules. 
Distribution. California; Canada. 
220 
