specimens now seen from California, 
Colorado (fig. 570), New York, and 
Utah is identical but differs 
considerably from that previously 
accepted as £. (^. ) inconspicua by 
Nowakowski and Spencer. In his 
monograph of this genus, Nowakowski 
treats C. inconspicua as one of five 
species under the £. ( P. ) atra 
superspecies. Differences are slight, 
but if this splitting is accepted, 
then it seems probable that the 
Canadian and European form represents 
another undescribed species in this 
complex. 
Cerodontha ( Poemyza ) macminni Spencer, 
new species 
(Figs. 565, 566) 
Head. Frons almost 1.5 times width of 
eye, not projecting above eye in 
profile; 2 ors, 2 ori; orbit consider- 
ably widening below level of lower 
ors; lunule high, narrow, upper margin 
at level of lower ors; gena narrow, at 
rear only 0.1 height of eye; 3d 
antennal segment small, round, arista 
long, only slightly pubescent, longer 
than wide, only slightly shorter than 
height of eye. 
Mesonotum. 3+1 dc or presutural 
lacking, acr in some 6 rows, prsc 
undeveloped . 
Wing. Length 2. 3-2.5 mm, C extending 
strongly to M 1+2, last section of 
M 3+4 almost 1.5 times length of 
penultimate. 
Color. Frons, antenna, and palpus 
black, lunule faintly grayish; orbit 
either uniformly black or narrowly 
yellowish along inner margin; 
mesonotum black, only weakly shining; 
upper margin and hindcorner of 
mesopleuron bright yellow, pleura 
otherwise black; legs with femora 
black but bright yellow distally for 
slightly more than femoral width, 
tibiae narrowly yellow adjoining 
femora, otherwise black, tarsi paler, 
yellowish brown; abdomen uniformly 
black, moderately shining; squama, 
including margin and fringe, yellow; 
wing with all veins moderately dark, 
brown. 
Male genitalia. Aedeagus (fig. 565) 
with distal tubules not diverging, 
with uniform ventral curvature and 
distinct dilation distally; sperm pump 
with small blade, only lightly pig- 
mented . 
Host/early stages. Carex crus-corvi . 
Larvae forming long, brownish blotch, 
up to 35 cm in length, with from 4 to 
16 larvae in single mine; pupation 
internal, with black puparia lying 
horizontal in leaf, completely unat- 
tached, posterior spiracles on common 
elongate extension, each with 3 bulbs, 
lower hooklike (fig. 566). 
Holotype m. , Mississippi, Washington 
Co., Experimental Forest, Leland, 
emerged 25.V.79 from puparia coll. 
V.ll from Carex crus-corvi growing in 
water-filled ditch beside road; 
paratypes 5m., 11 f., emerged 
25-28. V. 79, otherwise same data (G. 
McMinn). Holotype and paratypes in 
USNM, 4 paratypes in KAS. 
Remarks. This species is unique in 
being the only known Poemyza to feed 
on Carex . Externally it generally 
resembles C.. phragmitidis Nowakowski, 
a leaf miner on Phragmites in Europe, 
particularly in the form of the 
aedeagus (see Nowakowski, 1973: fig. 
122). The elongate extension of the 
puparium is also as in C . phragmitidis , 
but the arrangement of the posterior 
spiracles is different, with the two 
processes diverging from the common 
base and with the lower of the three 
bulbs conspicuously hooklike (fig. 
566). In this character it closely 
resembles C. alpina Nowakowski (see 
Nowakowski, 1973: fig. 230), belonging 
to Nowakowski' s muscina group. The 
jump in host from Poaceae to Cyper- 
aceae, which has occurred in this 
species, has thus been accompanied by 
evolutionary changes in the larva, 
including the feeding habit, and it 
therefore cannot be accurately included 
in any of the special groups into 
which Nowakowski divided this subgenus. 
278 
