faintly brownish between vti and upper 
ors; 3d antennal segment black, 1st 
and 2d slightly paler, yellowish; 
mesonotum black, moderately shining, 
with only small, indistinctly 
yellowish patches at hindcorners; 
humerus (apart from central black 
area), notopleuron, upper margin of 
mesopleuron, and wing base bright 
yellow; legs black, foreknee only 
faintly yellowish; squama yellow, 
margin and fringe black. 
Male genitalia. Aedeagus (figs. 906, 
907) with anterior pair of ventral 
sclerites flaplike, posterior pair 
short, basal sclerites partially 
fused, asymmetrical; sperm pump large, 
with broad blade. 
Host/early stages. Bidens , Helianthus , 
Rudbeckia . Larva forming large blotch 
mine (fig. 908); puparium brown, 
posterior spiracles each with 3 bulbs. 
Holotype m. , New York, Seneca Co., 
East Varick, ex Bidens frondosa , 
30.X. 56 (S. W. Frost); paratypes 
Illinois, Champaign Co., Urbana, 1 f . , 
20. VI. 15; Indiana, Tippecanoe Co., 
Lafayette, 1 m. , 27. V. (? year); South 
Dakota, Kingsbury Co., Erwin, VI. 08; 
(all J. M. Aldrich); Pennsylvania, 
Adams Co., Arendtsville, 1 f., ex 
Bidens frondosa , 5. VIII. 20; Adams Co., 
Colerain Park, 1 m. , ex Rudbeckia 
22. VII. 62; (both S. W. Frost); 
Virginia, Montgomery Co., Blacksburg, 
2,100 ft, 1 f., 28. V. 62 (J. G. 
Chillcott). Holotype in USNM, 
paratypes in USNM, CNC, and PSU. 
There are other specimens in poor 
condition in the S. W. Frost collection 
from Arendtsville, Pa., reared from 
Helianthus sp. Leaf mines have been 
seen from Maryland, Montgomery County, 
Cabin John Creek, 24. VIII. 76 and 
26.X. 78, and from Minnesota, Hennepin 
County, Minneapolis, Minnehaha Creek, 
10. IX. 78, all on Rudbeckia laciniata . 
Remarks. Frick (1956) misinterpreted 
this species as C^. allecta (Melander). 
C. allecta was described from St. 
Vincent, West Indies, and is a neo- 
tropical species, with confirmed 
records from Brazil, Dominica, 
Trinidad, and Venezuela (Spencer, 
1963a: 338; 1973b: 37; Spencer and 
Stegmaier, 1973: 182). This misinter- 
pretation was entirely understandable 
at the time, as the two species agree 
closely, particularly in the important 
character of the entirely yellow 
orbit, but C. allecta is somewhat 
smaller and slightly darker. The male 
genitalia of the two species are 
entirely distinct (figs. 906, 907, and 
Spencer, 1963a: figs. 52, a , 52, b ) . 
C. frickiana appears to be a more 
northern species, with no records 
south of Virginia and Pennsylvania. 
It can occur together with C. f lavi - 
notum, of which the known hosts are 
Arctium and Eupatorium . 
Calycomyza jucunda (Wulp) 
(Figs. 884-887) 
Agromyza jucunda Wulp, 1867: 161. 
Holotype male from Wisconsin in RNH. 
Phytobia coronata (Loew, 1869: 48), 
synonymy established by Frick, 1956: 
288. Lectotype male from Pennsyl- 
vania in MCZ designated here. 
Calycomyza jucunda , Steyskal, 1973a: 
191. 
This species has been confused with C. 
platyptera , feeding on Asteraceae, 
until its identity was established by 
Steyskal (1973a). In C . jucunda the 
mesonotum is distinctly less shining 
than in C. platyptera and there are 
also distinctive differences in the 
male genitalia. 
Essential characters of the aedeagus 
of C . jucunda are the short pair of 
anterior ventral sclerites, the fully 
fused, flaplike posterior sclerites, 
and the almost discrete basal 
sclerites, which have only a light 
membranous connection. The disti- 
phallus is not solidly black, as 
appears when using only direct light 
(Steyskal, 1973a: fig. 1, e) but 
distinctly paler distally when seen in 
reflected light (figs. 884, 885). The 
mesophallus is slender adjoining the 
300 
