Male genitalia. Aedeagus (fig. 1100) 
with distiphallus directed dorsally, 
divided (fig. 1101), supported basally 
by pair of broad sclerites that taper 
ventrally; paramesophalli directed 
forward, lying in same plane as basal 
sclerites, becoming membranous 
apically; sperm pump with large, 
rounded blade and narrow stalk. 
Host /early stages. Unknown. 
Holotype m. , Idaho, Latah Co., Moscow, 
no date (J. M. Aldrich), in USNM. 
Remarks. This species superficially 
resembles the dark form of P. ranunculi 
and was tentatively identified by Frick 
as P. ranunculi var. praecox . However, 
it is distinctly smaller, the costal 
ratio is less, and the male genitalia 
show that the two species are not 
related. 
Phytomyza arnicicola Lundquist 
(new to U.S.A. ) 
(Figs. 1247, 1365) 
Phytomyza arnicicola Lundquist. 1949: 
173. Holotype male from Swedish 
Lapland in ZIL. 
This species is host specific on 
Arnica and has hitherto been known 
from northern Sweden, Alaska, and 
Alberta. Leaf mines on Arnica 
chamissonis ssp. foliosa , possibly 
referable to P. arnicicola (fig. 
1365), were found in Nevada County, 
Calif. (Spencer, 1981: 457). 
Two specimens recorded here have been 
identified from Colorado. Although 
the aedeagus of these two specimens is 
virtually identical to that of the 
holotype (fig. 1247) and that 
illustrated by Griffiths (1974b: fig. 
5) from Alaska, the venation of the 
Colorado population differs in the 
ratio of the second to fourth costal 
sections, which is appreciably less 
than in the Swedish and Alaskan 
populations (although only single 
specimens have been seen from each 
area). In these the costal ratio is 
approximately 2.5, whereas in the two 
Colorado specimens it is 2.25 and 
1.5. With the limited material 
available, this difference can only be 
treated as local variation. 
It seems probable that P. arnicae 
Hering, 1925 (see Spencer, 1976a: 
384) , which is present in the 
mountains of central Europe and at 
Jasper, Alberta, in the Canadian 
Rockies (Griffiths, 1974b: 109), may 
also be present at high elevations in 
the United States. 
New records. 
Colorado, Lake Co., Independence Pass, 
11,100 ft, tundra, 1 m. , 31. VIII. 61 
(J. G. Chillcott); Larimer Co., 
Estes Park, 7,500 ft, 1 m. , 2. VII. 61 
(C. H. Mann); both in CNC. 
Phytomyza asterophaga Spencer (new 
to U.S.A.) 
(Figs. 1226-1228) 
Phytomyza asterophaga Spencer, 1969a: 
230. Holotype male from Alberta in 
CNC. 
This species has been known previously 
only from central Alberta, and 
according to Griffiths (1976b: 244) it 
is common around the city of Edmonton 
on its only known host. Aster 
conspicuus . A probable sister species, 
P. ciliolati Spencer (1969a: 234), 
feeds on other Aster species and may 
be found in the United States. Differ- 
ences between the two species are 
slight, with the notopleuron 
frequently somewhat paler in P. 
ciliolati and the distiphallus with 
even weaker pigmentation (Griffiths, 
1976b: fig. 4). 
New record. 
North Carolina, Macon Co., Highlands, 
3,000 ft, 5. VIII. 57 (J. G. 
Chillcott), in CNC. 
Phytomyza astotinensis Griffiths (new 
to U.S.A.) 
(Figs. 1238-1240) 
Phytomyza astotinensis Griffiths, 
1976b: 248. Holotype male from Lake 
Astotin, Alberta, in CNC. 
310 
