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University of California Publications in Zoology [Vol. 27 
cave in some individuals, convex in others; cranial crests well devel- 
oped, longitudinal segments beginning just posterior to external 
nares, extending posteriorly in parallel course or slightly divergent ; 
preorbital crest slight or wanting ; transverse postorbital crest usually 
well developed, but short; tympanic membrane oval, decidedly higher 
than wide, vertical diameter about two-thirds length of orbit ; parotoid 
gland close behind orbit, slender, about twice as long as wide, length 
nearly twice that of orbit, posterior ends of the two glands divergent ; 
chin acuminate oval or roundly oval in outline ; angle of jaw under 
tympanic membrane; vocal sac of male occupying entire chin region 
from margin of jaw posteriorly; about half of upper arm buried in 
skin ; forearm about equal to hand ; one large palmar tubercle, usually 
a smaller one at base of first digit, inner digit stoutest; digits in order 
of decreasing length, 3, 1, 4, 2; body slightly longer than broad; 
femur largely buried in skin of body; tibia equaling or slightly 
shorter than femur ; tarsus about two-fifths length of foot ; hind foot 
long, about three-fourths length of heacl-and-body ; metatarsal tubercles 
two, outer small and rounded, inner large with distinct free cutting 
edge ; toes small, in order of decreasing length, 4, 3, 5, 2, 1 ; web much 
restricted, edge slightly serrated. 
Tongue moderate, less than half width of mouth at angles of jaws, 
cuneiform, attached by anterior two-thirds to floor of mouth ; internal 
nares rounded, placed some distance back from margin of upper jaw. 
Surfaces generally rough; tip of snout and immediate margin of 
upper jaw smooth ; eyelids, dorsal and lateral surfaces of body, and 
exposed surfaces of limbs with many distinct conical, sharp-pointed 
warts up to 3 mm. in diameter, bases of adjacent warts sometimes 
confluent ; warts larger and sharper on females than on males ; 
plantar surface of hand and foot with numerous closely spaced smaller 
warts; ventral surface with fine even pattern of closely placed low 
conical or subconical warts, smallest and least rough on chin, largest 
and roughest at posterior end of body. 
General body coloration above dull yellowish brown ; narrow verte- 
bral stripe whitish ; larger warts; or groups of warts on back narrowly 
surrounded with black ; warts red-tipped ; limbs obscurely barred with 
black; undersurface of body plain dull yellow, sometimes with small 
irregular markings of black on breast ; tips of fingers and toes dark 
brown or blackish; larger tubercles on plantar surface of hand and 
foot dark yellowish brown ; metatarsal tubercles yellowish brown at 
base, black at tips; iris in life metallic yellow (color description partly 
adapted from Dickerson, 1906, p. 91). 
Stejneger (1890, p. 116) describes two “medium-sized” specimens 
from Tanner’s Gulch, 3 miles north of Tuba City, Painted Desert 
[= east-central Coconino County], Arizona, as follows: “Above pale 
olive-green with a somewhat lighter stripe down the middle of the 
back ; tip of tubercles pale red surrounded by black rings ; lower 
surface olive-white. ” Two other much smaller [yearling?] specimens 
taken at “Tanner’s Crossing of the Little Colorado” in the same 
vicinity are described in detail as follows : 
