246 University of California Publications in Zoology [Vol. 27 
Eana boylii, Dickerson (1906, pp. 221-222, pi. 2 [fig. 7], pi. 84 [figs. 264- 
266]). General account. 
Eana boylii, Van Denburgli (19125, p. 159). Occurrence in Oregon. 
Eana boylii boylii, Camp (19175, pp. 117-118, fig. 3). Critical; locality 
records. 
Eana boylii boylii , Grinnell and Camp (1917, p. 146, fig. 5). Range in 
California. 
Eana boylii boylii, Stejneger and Barbour, 1917, p. 36; 1923, p. 34). 
General range. 
Eana boylii, Boulenger (1920, pp. 469-471), part. Critical; locality rec- 
ords. 
Eana boylii boylii, Storer (1923, p. 8). In Coast Range. 
Eana boylii boylii, Grinnell and Storer (1924, pp. 663-665). In foothills 
of Yosemite region. 
Diagnosis. — Size small, head-and-body length up to 66 millimeters 
(2% inches) ; dorsal surface of body roughened with many fine 
pointed papillae ; vomerine teeth small, scattered on two elongate 
ridges between internal nares ; tympanic membrane like rest of head ; 
upper labial region without raised white ridge ; red not present in 
coloration of ventral surface; hind leg long, inside angle of bent 
tarsus reaching at least to nares and usually beyond ; each eyelid with 
light patch anteriorly and dark patch posteriorly. 
Comparisons — Distinguished from Scaphiopus by lack of cutting 
spade on hind foot and rounded instead of vertically elliptical pupil j 
from Bufonidae by absence of parotoid glands, and by presence of 
teeth in jaws ; from Hylidae, by larger size, lesser expansion of digits 
at tips and absence of dark stripe on side of head ; from Rana aurora 
ssp., and Rana pretiosa ssp., by rudimentary vomerine teeth, rough- 
ened tympanic membrane, absence of light stripe along upper lip, 
and (usually) absence of conspicuous dorsolateral folds; from Rana 
catesbeiama by smaller adult size, much smaller and less conspicuous 
tympanic membrane, and rougher dorsal surface; from Rana pipiens 
by broader form of body, rudimentary vomerine teeth, smaller and 
roughened tympanic membrane, absence of dorsolateral folds and 
presence of roughened body skin with ill-defined dark markings ; from 
Rana boylii muscosa by presence of light patch in front of dark area 
on upper eyelid, by darker general coloration, and less expanded tips 
of toes ; from Rama boylii sierras by longer hind legs, greater width of 
head, rougher tympanic membrane, absence of dorsolateral folds, and 
presence of light patch on upper eyelid. (Compare pi. 12, figs. 34, 36; 
pi. 17, fig. 51 [left].) 
Description. — Form stout, body broad, limbs short ; head broad yet 
pointed, oval in outline from above, thick in profile, muzzle descend- 
ing, rounded at tip ; external nares not terminal but nearer tip of 
snout than orbit ; internarial width greater than interorbital width ; 
canthus rostralis distinct, beginning at nostril, its length shorter than 
that of orbit ; orbit large ; interorbital space about two-thirds length 
of orbit; tympanic membrane rounded, smaller than, and separated 
slightly from, orbit, depressed below adjacent surface of head ; angle 
of jaw under tympanic membrane; lower jaw oval in outline; fore 
limb stout ; free portion of upper arm shorter than forearm ; hand 
longer than forearm ; fingers rather stout, blunt-ended, third longest, 
