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University of California Publications in Zoology [Vol. 27 
mens listed from Fort Walla Walla, Washington (Cope, 1889, p. 434). 
Two specimens from Humpeg Falls, Columbia County, and Butte 
Creek, Blue Mountains, in eastern Washington (nos. 5586, 5587, Mus. 
Vert. Zool.), are referable to this subspecies. The line of separation 
between luteiventris and pretiosa remains to be determined. 
Life-history. — Thompson (1913, pp. 54-55) says that in north- 
eastern Nevada this frog was common in irrigation ditches. 
It is quite aquatic, although the stomach contents, consisting of ants and water 
insects, indicate that at least part of the hunting is done on land. A few young 
specimens were taken a short distance from water on the banks of Anne Creek, 
but the adults were usually found along the edges of swiftly flowing streams or 
with only the head projecting from the vegetation of stagnant pools. When dis- 
turbed, they slipped quietly under the surface, but quickly reappeared, usually 
in the same place. Although they were almost as common as Eana pipiens, the 
two species were seldom found in the same pond. 
The larva described above was collected on July 10, 1912, on which 
date, according to the description given by Thompson, the dorsolateral 
folds were in evidence and the hind legs were well developed ; this 
would suggest a breeding season in March or April. 
Introduced Species 
Rana catesbeiana Shaw. Eastern Bullfrog 
(PI. 18, figs. 53-56; text fig. DD) 
Eana catesbeiana Shaw (1802, pp. 106-108, pi. 33). Original description, 
type from (eastern) North America [= Carolina?: see Stejneger and 
Barbour, 1917, p. 37.] 
Eana catesbeiana, Storer (1922, pp. 219-224, figs. 86, 87). Introduction 
and occurrence in California. 
Diagnosis. — Largest Salientian in California; head-and-body 
length up to 175 millimeters (7 inches) ; skin of back relatively 
smooth, without dorsolateral folds ; tympanic membrane equaling 
(females) or exceeding (males) diameter of orbit; a smooth, rounded 
ridge behind orbit and around dorsal and posterior margins of tym- 
panic membrane ; tympanic membrane of females concolor with body, 
of males dark brown marginally, with raised central portion of same 
color as body ; no red anywhere on under surface of body. 
Comparisons. — Distinguished from all other California Salientia 
by larger tympanic membrane ; from other species of Rana by larger 
adult size, relatively smooth upper surface, and by absence of dorso- 
lateral folds ; from Rama aurora ssp. and Rana pretiosa ssp. by absence 
of red in coloration. 
