The Philippine Journal of Science, C. Botany. 
Vol. XI, No. 1, January, 1916. 
THE GENUS LOXOGRAMME 
By Edwin Bingham Copeland 
( From the College of Agriculture , University of the Philippines, 
Los Banos. P. I.) 
four plates 
Loxogrctmme, as the name of a group of ferns, originated with 
Blume 1 who used it for a subgenus of Antrophyum. In dis- 
cussing the affinity (a curious expression in pre-Darwinian 
science) of Antrophyum, he points out a resemblance to Gram- 
mitis, especially to the species (now known as Polypodium) with 
coriaceus fronds and the sori oblique to the costa, and continues : 
“Plures adeo Grammitides ut sectionem propriam, a soris obliquis 
nomine Loxo gramme (a A 6£os obliquus et y pafJL/JLr) linea) insig- 
nitam, subjungimus Antrophyis.” 
As a generic name, Loxogramme dates from Presl, 2 with the 
following diagnosis. “Venae internae, tenuissimae, ramosis- 
simae, venulisque in maculas hexagonoideas elongatas inaequales 
anastomostantes et reticulam laxam efficientes. Sori dorso venae 
lateralis longioris unius aut duarum supra-positarum inserti, 
lineares, elangati, crassi, obtusi.” He adds that the rhizome is 
creeping; the fronds coriaceous, simple and entire, and the sori 
immersed, in the upper part of the frond. 
The first species listed by Blume is Antrophyum lanceolatum, 
the type of which is Grammitis lanceolata Swtz. It is partic- 
ularly fortunate that Swartz figured this fern himself 3 for 
Schkuhr, to whom we turn for figures of many illustrations of 
Swartz’s species, has in this case figured a diffenent fern, not 
belonging in the same genus, under this name. Blume in turn 
gives an excellent plate of his fern, which is still another species, 
but this time at least a Loxogramme. Presl also presents a 
figure, to illustrate the generic character (1. c. Tab. IX, Fig. 8), 
and labels it L. lanceolata; but it is the Antrophyum lanceolatum 
of Blume, not the Grammatis lanceolata of Swartz. 
The first species listed by Presl is Loxogramme coriacea 
[Grammitis coriacea Kaulf. in Spreng. Syst. Veg. 4 (1827) 71]. 
1 Flora Javae 2 (1828) 73. 
2 Tentamen Pteridographiae (1836) 214. 
8 Synopsis Filicum. Plate I, fig. J. 
43 
