528 The Philippine Journal of Science 1920 
united into a compound panicle. Long slender vasa efferentia 
convey the sperm to the region immediately anterior to the ace- 
tabulum, where they unite to form a single vas deferens. Where 
the short vas deferens enters the cirrus sac it bends to the left 
and enlarges to form the seminal vesicle. This organ twists on 
itself and then turns anteriad to form the penis. The latter 
is quite muscular and has the shape of a bottle with a long 
neck. The genital pore lies just behind the pharynx in the 
median line. 
The ovary is composed of numerous branches lying to the 
left of the ootype. It is relatively small. The vitellaria extend 
from the region of the acetabulum to the subdistal margin of the 
worm. The ventral follicles are strictly lateral, but the dorsal 
portion of the glands extends mesad in the posterior half of the 
worm. The vitelline ducts are less readily made out than in 
Fasciola hepatica. The transverse duct lies just behind the 
ootype. At its middle a short duct runs into the latter organ. 
The uterus coils on itself several times, then crosses over to the 
acetabulum and reaches the genital pore on the left of the 
cirrus sac. 
The eggs measure 140 to 160 y by 75 to 90 y , with an average 
of 145 y by 85 y, as distinguished from the measurements of 
Fasciola hepatica eggs, which have an average of 132 y by 70 y. 
DISCUSSION 
The literature on the genus Fasciola is considerable, even when 
one takes into account only the species closely related to Fasciola 
hepatica. With the removal of the species magna to a new 
genus, Fascioloides (Ward 1917 :3) (4), one is confronted only 
with the possible species F. hepatica, F. angusta Railliet, F. 
aegyptiaca Looss, and F. gigantica. A close inspection shows 
that the latter three species are alike in all respects in which they 
differ from F. hepatica. They are elongated flukes with parallel 
lateral margins. Their anterior cone is distinct but short. 
Their testes are richly branched but short and occupy a field 
much more restricted than those of F. hepatica. Ovary and 
vitellaria are richly branched. The lateral divisions of the gut 
are more numerous. The acetabulum is relatively large and 
close to the oral sucker. The cirrus pouch is small and much 
more contracted than in F. hepatica. Moreover the egg measure- 
ments are remarkably similar, averaging 147 /x by 82 /x for 
F. angusta, 150 y by 80 y for F. gigantica of Egypt, and 145 
a by 85 /x for the material which I have studied. 
