6 
COPELAND. 
of at most 34 known species, cosmopolitan, but with a boreal center of 
distribution. 
Pi mice flabellate rather than pinnate. 
Sterile and fertile segments separating just below sterile blade 1. B. lunar ia 
Common stalk very short 2. B. simplex 
Fronds at least tripinnatifid. 
Fertile segment arising below the base of the sterile. 
Sterile frond fleshy, typically with a hyaline margin. 
Segments widened on both sides above base 3. B. matricari.ae 
Segments not widened on both sides 4. B. obliquum 
Sterile frond without hyaline margin. 
Frond fleshy 5. B. ternatum 
Frond thin, herbaceous. 
Fertile segment arising above middle of the petiole 6. B. daucifplium 
Fertile segment arising lower down 7. B. japonicuin 
Fertile segment arising from base of sterile : 8. B. virginianum 
Fertile segment arising from racliis of sterile 9. B. lamiginosum 
1. Botrychium lunaria (L.) Swtz. 
Stipe stout, fleshy, 3 to 10 cm high; sterile segment 2 to 6 cm long, 
about 2 cm broad, pinnate, rachis flattened rather than winged; pinnae 
usually equal, broader than long, sessile or nearly so, subcuneate, entire 
or variously incised on the outer side, fleshy, glabrous, the lowest pair 
sometimes enlarged, and doubly incised; fertile segment arising from 
(or just below) the base of the sterile, its stalk 3 to 7 enr long, the 
blade (panicle) lanceolate deltoid, pinnae stalked, the lowest pair pinnate. 
Himalayas to Europe and Siberia; North America, Patagonia, New Zealand, 
southern Australia. 
2. Botrychium simplex Hitclic. 
Like B. Lunaria, but decidedly smaller and more simple, and the stalks 
of the two segments separating well below the base of the sterile one. 
Japan, North America, northern Europe. 
3. Botrychium matricariae (Schrank) Spr. 
Stipe separating less than halfway up to the base of the sterile 
segment; young frond hairy; sterile blade deltoid, in Chinese specimens 
5 cm long, 4 cm broad, tripinnate, the lobes small, oval, widening abruptly 
on both sides above the base, or even cordate, margin hyaline because of 
thick epidermis; veins spreading above the base; stipe of the ample fertile 
segment 10 cm long. The plant grows larger elsewhere. 
Central China, northern Asia, Europe and America. 
4. Botrychium obliquum Miihl. 
Like the preceding, but the common stipe very short — only 1 cm long 
in Chinese specimens — the sterile segments broader than long, its ulti- 
mate divisions more ample (i. e., less cut), and oblique, usually not much 
widened above the base on the upper side, and the veins less spreading. 
Su-Chuen; eastern North America. 
