284 
COPELAND. 
Gunong Bengkarum, altitude 1,050 m, leg. 0. J. Brooks, No. Ifi : growing in and 
beside a very cold stream. 
This little fern is unmistakably related to Saccoloma moluccanum (3B1. ) Mett., 
and to Lindsaya cultrata (Willd.) Sw. The pinnae strongly resemble the 
ultimate pinnules of S. moluccanum, and the sori are alike except in position. 
The rhizome and the aspect of the plant are like the Lindsaya. The origin of 
Lindsaya, Schizoloma and Odontosoria has always been a mystery, to which this 
fern offers a very interesting clue. I by no means believe that P. Brooksii is 
itself the ancestor of Lindsaya or Odontosoria or Saccoloma; but it presents so 
interesting a combination of the characteristics their common ancestors must 
have had, that it is reasonable to suppose that it has retained with less modifica- 
tion the less specialized, more generalized character of the more primitive and 
now unknown real ancestor. 
In the same connection it may be observed that S. moluccanum is with doubtful 
propriety referred to Saccoloma. It has had at least four specific names in 
Microlepia, to which its affinity is clear. 
Schizoloma heterophyllum (Dry.) J. Sm. var. Speluncae Copel. var. nova. 
Stipite 4-10 cm alto nisi versus basin castaneam viride; fronde ca. 5 
cm alta; pinnis maximis 33 mm longis,. 12 mm latis, obtusis, basibus late 
cuneatis fere aequilateralibus, integris vel rarissime subincisis, membra- 
naceis; venulis laxe anastomosantibus ; soris vix marginalibus, continuis. 
Sandakan, in a cave at base of sandstone cliff, facing the sea, Foxworthy 578. 
Except for the suspicion that the peculiar environment may. be responsible for 
the thin, broad, entire pinnse, I would not hesitate to describe this as a new 
species. 
Asplenium trifoliatum Copel. spec. nova. 
Euasplenium foliis maximis, rliizomate suberecto, 7 mm crasso, lignoso, 
radices validas multas emittente, dense paleaceo, paleis membranaceis 
ovato-lanceolatis integris acuminatis 6 mm longis brunneis; stipitibus 
15-20 cm altis, confertis, paleaceis, paleis sursum minoribus et subdeci- 
duis ; fronde pinnata ; pinna utroque latere una, decurr'enti-adnata, ca. 
12 cm longa, 3-5 cm lata, eaudata; pinna terminale 40 ad 50 cm alta, 
fere 10 cm lata, eaudata, argute serrata, lierbacea, supra glabra, atro- 
viride, infra pallidiore, et ad costam atro-brunneam prominentem et 
sparsissime squamulis minutis ad laminam paleaqea ; soris laete brunneis, 
inaequalibus, costa et margine remotis, vix 1 mm latis, indusio per-, 
sistente, sporis cornutis. 
Sambas, near Tringos, on rocks in low-lying jungle, leg. C. J. Brooks, No. 26 1 
In some respects this species strongly suggests A. epiphyticwin Copel., of 
Mindanao, which I regard as a probable source of the genus Stenochlaena.' It 
has also enough characteristics in common with A. squamulatum to make their 
affinity very probable. ' It may, therefore, well represent the connection between 
Thamnopteris and the larger and probably older group, Eu-asplenium. On the 
two fronds sent me I find only two pairs of veins anastomosing at the margin. 
The veins end uniformly in elongate hydathodes. 
