210 
BROWN. 
According to Schaar,^ Unger/ in a paper which is not avail- 
able to the writer, says in reference to Rafflesia patma, “Wir 
sehen den Parasiten mit der Nahrpflanze in der Art vereinigt, 
dass derselbe unmittelbar iiber dem Holzkorper seines Tragers 
entspringt und durch sein Gefasssysten mit dem Gefasssystem 
der Nahrpflanze anastomosirt.” 
Solms-Laubach ® showed that the vegetative portion of Pilo- 
styles haussknechtii, one of the Rafflesiaceae, consisted of simple 
cell strands in the pith, medullary rays, vascular elements and 
bark of species of Astragalus. These cell-strands Solms-Laubach 
designated as mycelium on account of their resemblance to the 
mycelium of a fungus. They extend into the bases of the leaves 
where they form cushions from which the flower buds are de- 
veloped. These buds break through the tissues of the host and 
appear upon the basal portion of the leaves. 
A similar structure of the vegetative portion of the plant has 
been shown by Solms-Laubach ® and Peirce ^ for Brugmansia 
zippelii, by Solms-Laubach ® and Pierce ^ for Rafflesia patma and 
by Schaar ° for Rafflesia rochussenii. 
The present study is based upon material of Rafflesia manil- 
lana Teschem.^° collected during January and February, 1912, 
on a forested ridge of Mount Maquiling, Province of Laguna, 
Luzon, at an elevation of about 400 meters. 
The original description of Rafflesia manillana by Tesche- 
macher was based on small unopened male buds, preserved in 
alcohol. The only description of an expanded flower is a short 
and unsatisfactory one by Blanco of a male flower of R. phil- 
’ Schaar, F. Uber den Ban des Thallus von Rafflesia Rochussenii Teysm. 
[und] Binn. Siztber. Acad. Wien 107 (1898) Abt. 1: 1039-1056. 
* Unger, F. Beitrage zur Kenntniss der parasitischen Pflanzen. Annalen 
des Wiener Museums der Naturgeschichte. 2 (1840). 
Solms-Laubach, H. Graf zu, Uber den Thallus von Pilostyles haus- 
sknechtii. Bot. Zeit. 32 (1874) 49-59, 65-74. 
® Solms-Laubach, H. Graf zu. Die Entwicklung der Bluten bei Brug- 
mansia zippelii und Aristolochia clematitis. Bot. Zeit. 34 (1876) 449. 
’ Peirce, G. J. On the Structure of the Haustoria of some Phanerogamic 
Parasites. Ann. Bot. 7 (1893) 291-327. 
* Solms-Laubach, H. Graf zu, Rafflesiaceae. Engler & Prantl Nat. 
Pflanzenfam. 3^ (1889). 
“Schaar, F. Sitzber. Acad. Wien 107 (1898) Abt. 1: 1039-1056. 
“ Teschemacher, J. E. On a new species of Rafflesia from Manilla. 
Boston Journ. Nat. Hist. 4 (1844) 63-65, t. 6; Ann. & Mag. Nat. Hist. 9: 
381-384. 
“Blanco, M. Flora de Filipinas, ed. 2 (1845) 565. 
