290 
COPELAND. 
The greatly enlarged lowest pair of pinnae and the strong development of the 
basiscopic pinnules of this pair are unique in Athyrium, and strongly suggest 
that the plant is really a Dryopteris. 
GROUP OF A. .1 A PON 1 COAL 
3. A. japonicum (Thunb.) Copel. comb. nova. Asplenium japonicum Tliun- 
berg FI. Jap. (1784) 334; Diplazium, Beddome; Athyrium Copelandi Christ in 
Philip. Journ. Sci. 2 (1907) Bot. 161. 
Batanes Islands, Bur. Sci. 3812 Fenix; Babuyan, Bur. Sci. 3899 Fenix ; Mount 
Data, Copeland 1909 ; Mount San Cristobal, Laguna, Copeland s. n. : Mindoro, 
Merrill 5920. 
Japan to India. 
Even though construed fairly narrowly, excluding such forms as Diplazium 
Petersenii, this is still a decidedly variable fern, and I can not, judging from the 
single collection of A. Copelandi, see sufficient differences to justify holding it 
distinct. 
4. Athyrium grammitoides (Presl) Milde. Diplazium grammitoides Presl 
Epim. (1849) 84. Asplenium brachypodum Baker, Syn. Fil. (1874) 490. 
Asplenium toppingianum Copel. in Perkins Fragmenta (1905) 184. Athyrium 
nanum Christ in Philip. Journ. Sci. 2 Bot. (1907) 161. 
“Philippines” Cuming 56: Batilao, Bontoc, Copeland 1896; Benguet, Topping 
191/, 281/, 309, Elmer 5787, 6285, Copeland 1801, Bur. Sci. 2872 M earns ; Mount 
Mariveles, Copeland 201/1; Cavite Province, Bur. Sci. 1309 Mangubat ; Mount 
Maquiling, Copeland s. n. ; Tayabas, Elmer 801/1: Mindoro, Merrill 5921: Mount 
Canlaon, Negros, Copeland 2077 : Lanao district, Mindanao, Clemens 656, 1125a ; 
Zamboanga, Copeland llfila; Mount Apo, Copeland 11/91, Williams 2519. 
A decidedly variable little species, but, as I here' construe it, clearly distinct 
from all others known to me except A. japonicum. Forms are rarely found 
which seem as near the latter as to A. grammitoides. A. Toppingii is what I 
now believe to be typical grammitoides. The variety mistum is a form occa- 
sionally found wherever the species is, and not entitled to consideration as a 
stable variety. 
One of our sheets of Cuming’s No. 56 is entirely A. grammitoides, but the 
other contains also an immature plant which may be A. Williamsi. Judging 
from Baker’s description I think he may have had the mixture before him, but 
his diagnosis applies most nearly to small specimens of .4. grammitoides. A. 
nanum is described as a dwarf, such as is occasionally found in various places; 
but the type collection of .4. nanum, Clemens 656, includes also the typical form 
and the still large mistum. 
5. Diplazium Acrotis Christ in Bull. Herb. Boiss. II. 6 (1906) 1000. 
Dr. Christ distinguishes this little fern from A. grammitoides by the absence 
of a wing on the almost filiform rachis, very regular pinnae, each with a sharp 
rectangular auricle, and paleae 5 mm long on the stipe. 
GROUP OF A. FILIX-FOEMINA. 
6. Athyrium drepanopteron (Ivze.) Moore. 
Benguet, Topping 237, 21/9, 333, Pond s. n., Elmer 61/98, Copeland 1800, Boyce 
s. «., For. Bur. 5090 Curran, Bur. Sci. 2751/, 31/35, 31/36, 3519, 1/191, 1/515, 
1/521, it 533 M earns. 
Japan to northern India. 
