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COPELAND. 
true Goniophlebium. They have also a general aspect familiar in Goniophlebium, 
but still more so in Eupolypodiuni. In Goniophlebium simply pinnatifid fronds 
with the segments dilated at the base are much commoner in the American plants 
referred to this sub-genus, but occur also in a number of Old World species, 
especially in northern India. On geographical grounds, and because of the 
resemblance to Eupolypodiuni, and because of less structural specialization, this 
would naturally be regarded as the primitive form of Goniophlebium, from which 
“Shellolepis” is a derived group. 
The primitive and generalized character of these plants is much more evident 
in the venation. In both species there is a costal row of areolae, but these areolae 
contain no free veins. The sori are borne sometimes on the vein which incloses 
the areola, sometimes against this vein but on a rudimentary branch excurrent 
from it. P. proavitum occasionally has other veins excurrent from this one. 
The venation then is neither that of typical Goniophlebium nor of Phymatodes, 
but is a generalized one, suggesting both of these. The structural similarity 
of Goniophlebium (Shellolepis) and Phymatodes and Selliguea I have already 
noted. 1 Mr. Maxon has since told me that in his opinion these constitute a single 
group properly to be regarded as a genus. The discovery of these Bornean ferns 
is the best possible support for liis judgment as to the affinity of Phymatodes 
(and Selliguea) to Goniophlebium, and shows that Goniophlebium, including 
them would certainly be a natural genus : but, so is Polypodium natural ; and, 
unwieldy though it is, there is no hurry about dismembering it. 
Polypodium sablanianum Christ Philip. Journ. Sci. 2 (1907) Bot. 177. 
Sarawak, Mount Bidi, leg. Brooks. 
Fronclibus pubescentibus, grandibus, herbaeeis, seriebus areolarum 
praestantium duabus, a P. myriocarpo (Pr.) Mett. diversum. 
The fronds of the Bornean specimens are more than 60 cm long and about 6 
cm broad. 
Described from Philippine specimens. 
Polypodium ceratophyllum Copel. spec. nova. (Plate VII.) 
Polypodium epipliytieum, rhizomate repente, 1 mm crasso, paleis 
ferrugineis 2.5 mm longis setiformibus basibus peltatis vestito; stipite 
gracile, brunneo-fulvo, articulato, ca. 8 cm alto sed supra medium lamina 
decurrente alato; fronde, ala decurrente exclusa, ca. 2.5 cm alta, ca. 4 
cm lata, iterum bifurcata, ramis late divergentibus, superioribus ca. 2 
mm latis, obtusis, coriaceis, glabris, margine angusta cartilaginea sub 
lente levissime ineisa; venis occultis, laxe anastomosantibus ; soris super- 
ficialibus, utroque latere costae nigrae unfseriatis. 
Sarawak, Mount Poe, alt. 1,300 m, Foxworthy 205. 
A species of the Microterus group, but with dichotomous fronds. On a sepa- 
rate rhizome are sterile fronds, broad and short and only once or twice forked; 
they may be fronds of an immature plant, or the sterile fronds of an adult. 
Syngramma angusta Copel. spec. nova. 
Ehizomate repente, lignoso, 2.5 mm crasso, paleis lanceolatis minutis 
obscuris coronato : stipitibus c-onfertis, nisi ad baseos sparse paleaceas 
glabris, frondium sterilum 5-8 cm fertilium ca. 20 cm altis; fronde 
This Journal, 2 (1907) Bot. 74. 
