MICROLEPIDOPTERA OF NEW GUINEA 
99 
Sigi Camp, 1500 m, February 15, 1939 (allotype), February 26, 1939 
(holotype). Two specimens. Allied to I. melanoclera Meyrick, but much 
smaller and with transverse fascia complete. 
CACOECIINI 
“Group A” Meyrick (partim), in Wytsman, Gener. Ins., vol. 149, p. 2 anti 7, 1913. 
Archipsidii Pierce & Metcalfe, Genit. Brit. Tortr., p. 1, 1922. 
Cacoecidii Diakonoff, Zool. Meded. Mus. Leiden, vol. 21, p. 151, 1939. 
Cacoeciades Diakonoff, Treubia vol. 18, p. 31, 1941. 
This tribe represents by far the greater part of the Tortricinae in the 
present collection and also apparently in the fauna of New Guinea. The 
following six genera belong to a natural group, and, being related with 
Isotenes, form a connection with the previous tribe. They are as a rule 
obscurely greyish-coloured species with a more or less ciliated costa in 
the fore wing, and often with veins 7 and 8 separate. Their resemblance 
to Isotenes is deceiving, but a thorough study of the facies, of the shape 
of the fore w r ing, and especially of the clear cut and large difference of the 
genitalia (of the two sexes) lender characteristics for their separation from 
the Zacoriscini. The soeius in Leptochroptila is of the zacoriscid type, but a 
corethrogyne is entirely absent. Upon that genus follows Anisotenes 
gen. nov., a relative of Isotenes, but without any traces of a corethrogyne, 
and with a distinctly developed, projecting and separated but small soeius 
of the cacoeciid type, instead of a single row of bristles along the rim of the 
sclerotized ventral parts of the uncus (above the articulations of the 
gnathos), as in the Zacoriscini. In other, less specialised genera, like 
Isochorista and Parachorista, the socii are considerably developed and 
form moderate or large appendages, densely covered with fine bristles. 
Thrincophora stands rather apart from the other genera. 
In our opinion the soeius is an important part of the male genitalia, 
being with a few exceptions an indicator of the stage of development, not 
only in this tribe but throughout the subfamily Tortricinae. The soeius 
acquires highest development in the relatives of Peronea, and shows a 
gradual reduction throughout the Cnephasiini towards the Cacoeciini to 
disappear altogether as a separate protuberance in the Zacoriscini. 
The two genera following upon Thrincophora, viz., Chresmarcha Meyrick 
and Adoxophyes Meyrick, form a rather less apparent entity, super- 
ficially. They possess vein 3 in the fore wing originating from before angle, 
while veins 7 and 8 are mostly long-stalked. The soeius is moderate, 
pending, the gnathos is strong; the cestum is absent. Adoxophyes shows the 
tropical cacoeiid colouring and markings, but Ghresmarcha forms a striking 
exception : it is a perfect mimic, inter alia, of the Zacoriscine genus Chiono- 
thremma, and without doubt has diurnal life habits. 
The third group comprises Cacoecia and Ilomona and their relatives, 
with veins 7 and 8 in the fore wing mostly stalked, while vein 3 originates 
from the angle. The uncus is narrow and strongly curved. The soeius is 
