102 
MICROLEPIDOPTERA OF NEW GUINEA 
because it forms a connection between the two tribes : the Zacoriscini and 
the Cacoeciini, as is already said above. Therefore it is necessary to submit 
this genus to a thorough discussion. 
Anisotenes is closely allied to Isotenes Meyrick 1938, and the resem- 
blance of the facies of these two genera is very great. However, the former 
does not possess the typical characteristics of the tribe Zacoriscini; it is a 
true Cacoeciid. The differences between the two genera can be outlined 
as follows. The females often possess a similarly ciliated costa, and both 
fore and hind wing have a very similar shape and neuration. (The neuration 
in Anisotenes is rather variable as to the origin of veins 3 — 5 and 6 — 7 in 
hind wing.) The two genera can be separated at once by the entire absence 
of a corethrogyne in the last mentioned genus, where the anal abdominal 
segment is smoothly and evenly scaled, and only the eighth segment has 
a small thin marginal dorsal collar of scales, forming a modest anal tuft 
of the ordinary type, present in all the Tortricidae. The seventh ventrite 
is normally foimed, and after maceration shows no traces of expansible 
plicate membranes, (so characteristic of all female members of the Zaco- 
riscini) upon which the dense scaling of the corethrogyne is implanted. 
The males of Anisotenes may be separated from those of Isotenes by a 
broader wing which is gradually narrowed towards the base, while the 
costal edge is mostly rather smooth. The fore wing is Isotenes is smaller and 
rather abruptly narrowed towards the base, which is a typical feature; 
the folded part of the costa is little curved but considerably oblique. The 
genitalia in Anisotenes possess a rather small but distinctly separated 
digitiform pending socius, which part in Zacorisca is reduced to a series of 
bristles above the articulations of the arms of the gnathos. The great 
importance of the development of the socius for the separation of sub- 
families was already brought forward in the discussion of the present 
tribe. A single species of Isotenes ( mesonephda ) has a minute verrucose 
protuberance at the place where in other subfamilies a well-developed 
socius originates. Furthermore, in most species of Isotenes the costal patch 
in the fore wing is formed by two costal quadrate dots, mostly dark grey, 
and a third elongate-rectangular fuscous or fulvous spot obliquely below 
the costal dots, closer under the first of these. This characteristic is small 
but invariably present in all but one species (not distinct in pudens, of 
which we possess a single, somewhat damaged specimen). The costal patch 
in Anisotenes mostly is an equilateral triangle, and is often marked with 
three dots at its angles (as in Isotenes), but in A nisotenes the three dots are 
of equal shape, size and colour. 
Structurally Anisotenes is almost congruent with Harmologa Meyrick, 
1883; therefore we hesitated for a long time, whether the following species 
from New Guinea belong to that genus or not. At present, however, we 
are satisfied that this resemblance is oidy superficial as the type of the 
neuration in the type of the genus, Harmologa oblongana (Walker), 1863, 
from New Zealand, is quite different: it possesses a broad cell which is 
