MICROLEPIDOPTERA OF NEW GUINEA 
87 
EUCOSMINAE 
(Olethreutinae auctorum ) 
The Eucosminae are far less numerous in New Guinea than the Tor- 
tricinae, viz., in the proportion of one to three. This fact and its signi- 
ficance were already discussed in part I (pp. 29—30). It may he repeated 
here that we adopted Heinrich’s view that the Eucosminae must have 
developed out of the Tortricinae, and not on the contrary. 
In the near future we plan to revise the Eucosminae from the western 
part of the Malay Archipelago, and at that occasion we propose to discuss 
to some extent the general aspects of the taxonomy of this subfamily. At 
present we shall confine ourselves to a few remarks. 
In connection with the discussion of the distribution of the Eucosminae 
in the Papuan region as dealt with on p. 29 of part I, we may remark here 
that the subfamily seems to have obtained a firm foothold in New Guinea. 
Though this subfamily is not very extensively represented in that island, 
several endemic genera have developed here. The genus Hermenias 
Meyrick, also known from India, Ceylon, and Australia, presents a long 
series of closely allied species from Central New Guinea and in that 
country apparently is in a stage of rich development. 
Heinrich’s division of the family in three subfamilies is adopted by us. 
We prefer, however, provisionally and tentatively, to adhere not to his 
nomenclature but to that of T. Bainbrigge Fletcher as laid down in 
the latter’s ‘'Generic names used in Microlepidoplera” , 1929. Consequently 
we use the name Argyroplocini instead of Heinrich’s Olethreutinae, and 
Enarmoniini instead of Laspeyresiinae, on the strength of the priority of 
the respective generic names: Argyploce Hubner 1826 ( Verz ., p. 379, 
genotype arbulella Linne) supercedes Olethreutes Walsincham 1900 {Ann. 
Mag. Nat. Hist., ser. 7, vol. 6, pp. 127—130, genotype arcuella Clerck), 
Olethreutes Hubner 1806 ( Tentamen , p. 2) is invalid (cf. Opinion 97 of the 
International Rules of Zoological Nomenclature); Enarmonia Hubner 
1826 (Verz., p. 375, genotype woeberiana Schiffner) supercedes Laspey- 
resia Hubner (tom. cit., p. 381, genotype corollana Hubner). 
As far as concerns the phytogeny of the three subfamilies, Heinrich 
regards the Olethreutinae as being the most primitive, from which both 
the Laspeyresiinae and the Eucosminae have developed. This opinion 
finds little support in the characters of the male genitalia, the enarmoniine 
(= laspeyresiine) type showing close affinities to the peroneine group in the 
Tortricidae and also to certain Schoenotenidae; this type less closely 
approaches the male genitalia in the subfamily Choreutinae of the "family 
Glyphipterygidae, from which all tortricoid families may have developed. 
Consequently for the time being we are inclined to regard the Enarmoniini 
as the oldest group of the Eucosminae; the Enarmoniini link this sub- 
family with the (ancestral) Tortricinae (see also part I, pp. 29 — 30). 
The tribe Eucosmini is possibly not as highly specialised as has been 
