134 
Fishery Bulletin 108(2) 
Research (CIFAR). We thank V. Gallucci, J. Rice, A. 
Andrews, and W. Strasberger for field and laboratory 
assistance, and G. Bargmann, S. Rosen, and J. Topping 
at the Washington Department of Fish and Wildlife 
for reading spines and training. We acknowledge the 
National Marine Fisheries Service; Alaska Department 
of Fish and Game; chartered vessels and crew of the 
FVs Kingfisher , Winter King, and Sea View, commercial 
fishermen in Yakutat, Cordova, and Kasilof; Gauvin 
and Associates, LLC., and Alaska Pacific and Trident 
Seafoods for kindly providing sampling opportunities. 
Finally, we are grateful to T. Quinn II and K. Goldman 
for considerable helpful analytical advice. 
Literature cited 
Araya, M., and L. A. Cubillos 
2006. Evidence of two-phase growth in elasmobranchs. En- 
viron. Biol. Fishes 77:293-300. 
Avsar, D. 
2001. Age, growth, reproduction and feeding of the 
spurdog ( Squalus acanthias Linnaeus, 1758) in the 
southeastern Black Sea. Estuar. Coast. Shelf Sci. 52: 
269-278. 
Beamish, R. J., and G. A. McFarlane. 
1985. Annulus development on the second dorsal spine 
of the spiny dogfish ( Squalus acanthias) and its valid- 
ity for age determination. Can. J. Fish. Aquat. Sci. 
42:1799-1805. 
Beverton, R. J. H., and S. J. Holt. 
1957. On the dynamics of exploited fish populations. U.K 
Ministry of Agriculture and Fisheries, Fisheries Inves- 
tigations 2, 533 p. 
Boldt, J. 
2003. Ecosystem considerations for 2004. In Stock 
assessment and fishery evaluation report for the ground- 
fish resources of the Bering Sea/Aleutian Islands and 
Gulf of Alaska region, 335 p. [Available from North 
Pacific Fishery Management Council, 605 W. 4th Ave., 
Suite 306, Anchorage, AK 99501.] 
Bonfil, R. 
2005. The purpose of stock assessment and the objectives 
of fisheries management. In Management techniques 
for elasmobranch fisheries ( J. A. Musick, and R. Bonfil, 
eds.), p. 6-14. FAO Fisheries Tech. Paper 474. 
Braccini, J. M., B. M. Gillanders, and T. I. Walker. 
2007. Comparison of deterministic growth models fitted 
to length-at-age of the piked spurdog ( Squalus mega- 
lops) in south-eastern Australia. Mar. Freshw. Res. 
58:24-33. 
Burnham, K. P., and D. R. Anderson. 
2004. Multimodel inference: understanding AIC and BIC 
in model selection. Soc. Meth. Res. 33:261—304. 
Cailliet, G. M., and K. J. Goldman. 
2004. Age determination and validation in chondrich- 
thyan fishes. In The biology of sharks and their rela- 
tives (J. F. Carrier, J. A. Musick, and M. R. Heithaus, 
eds.), p. 399-447. CRC Press, Boca Raton, FL. 
Cailliet, G. M., W. D. Smith, H. F. Mollet, and K. J. Goldman. 
2006. Age and growth studies of Chondrichthyan fishes: 
the need for consistency in terminology, verification, 
validation, and growth function fitting. Environ. Biol. 
Fishes 77:211-228. 
Campana, S. E. 
1990. How reliable are growth back-calculations based 
on otoliths? Can. J. Fish. Aquat. Sci. 47:2219-2227. 
2001. Accuracy, precision and quality control in age deter- 
mination, including a review of the use and abuse of age 
validation methods. J. Fish Biol. 59:197-242. 
Campana, S. E., C. Annand, and J. I. McMillan. 
1995. Graphical and statistical methods for determining 
the consistency of age determinations. Trans. Am. 
Fish. Soc. 124:131-138. 
Campana, S., C. Jones, G. A. McFarlane, and S. Myklevoll. 
2006. Bomb dating and age validation using the spines 
of spiny dogfish ( Squalus acanthias). Environ. Biol. 
Fishes 77:327-336. 
Carlson, J. K., and I. E. Baremore. 
2005. Growth dynamics of the spinner shark (Carcha- 
rhinus brevipinna) off the United States southeast and 
Gulf of Mexico coasts: a comparison of methods. Fish. 
Bull. 103:280_291. 
Compagno, L. J. V. 
1984. Sharks of the world: an annotated and illustrated 
catalogue of shark species known to date, pt. 2: Car- 
charhiniformes, 675 p. FAO Fish. Synop. 125 (vol. 
4). FAO, Rome. 
Courtney, D., C. A. Tribuzio, K. J. Goldman, and J. Rice. 
2006. GOA sharks. In Stock assessment and fishery 
evaluation report for the groundfish resources of the 
Gulf of Alaska for 2007, Appendix E, p. 481-561. [Avail- 
able from North Pacific Fishery Management Council, 
605 W. 4 th Ave., Suite 306, Anchorage, AK 99501.] 
Driggers, W. B., J. K. Carlson, D. Oakley, G. Ulrich, B. Cullum, 
and J. M. Dean. 
2004. Age and growth of the blacknose shark, Carcharhi- 
nus acronotus, in the western North Atlantic Ocean with 
comments on regional variation in growth. Environ. 
Biol. Fishes 71:171-178. 
Dudewicz, E. J., and S. A. Mishra. 
1988. Modern mathematical statistics, 864 p. John 
Wiley & Sons, Inc., New York, NY. 
Fabens, A. J. 
1965. Properties and fitting of the von Bertalanffy growth 
curve. Growth 29:265-289. 
Fahy, E. 
1989. The spurdog Squalus acanthias (L) fishery in 
south west Ireland. Irish Fish. Invest. Ser. B (Mar.) 
32, 22 p. 
Francis, R. I. C. C. 
1990. Back-calculation of fish length: a critical review. 
J. Fish Biol. 36:883-902. 
Goldman, K. J., S. Branstetter, and J. A. Musick. 
2006. A re-examination of the age and growth of sand 
tiger sharks, Carcharias Taurus, in the western North 
Atlantic: the importance of ageing protocols and use of 
multiple back-calculation techniques. Environ. Biol. 
Fishes 77:241-252. 
Gulland, J. A. 
1969. Manual of methods for fish stock assessment: part 
1, Fish population analysis. FAO manuals in fisheries 
science, no. 4, 154 p. FAO Rome. 
Hauser, L. 
2009. The molecular ecology of dogfish sharks (Squalus 
acanthias) . In Biology and management of dogfish 
sharks (V. F. Gallucci, G. McFarlane, G. Bargmann, 
eds.), p. 229-252. Am. Fish. Soc., Bethesda, MD. 
