Zimmermann: Maturity and fecundity of Atheresthes stomias 
605 
4.500 -| 
4 000 - 
3 500 - 
3.000 - 
2.500 - 
2.000 - 
1.500 - 
1.000 - 
0.500 - 
A 
o Early perinucleus 
□ Late perinucleus 
x Cortical alveoli 
* Early vitellogenesis 
a Late vitellogenesis 
* Migratory nucleus 
A 
A 
O 
O 
o 
o 
o 
o 
D go 
□“ v A** . ° 
□ A a a 
X A* X , 
A A * V * 
X * 
□ □ * A * > 
, B 0 £□* ° *□ °i X 
3 n « 8x X 
0D 
A 
S 0 000 
10 
20 
30 
40 
50 
60 
70 
80 
90 
cc 
Q. 
CD 
X 
2.5 n 
B 
□ Immature 
* Mature 
1.5 - 
1 - 
0 5 - 
0 -I 1 1 1 1 1 1 1 1 1 
10 15 20 25 30 35 40 45 50 55 
Length (cm) 
Figure 4 
Hepatosomatic index by length and maturity stage for female arrowtooth flounder (A) and for male 
arrowtooth flounder (B). 
with increasing female length (Fig. 4A). There was 
no difference in HSI between mature and immature 
males (unpaired f-test; df=27, P>0.05), and no trends 
in the data (Fig. 4B). 
Gonadosomatic index varied significantly among 
female maturity groups (single-factor AN OVA; df=5, 
F=91.9, PcO.OOl). The Tukey test revealed that de- 
spite more than doubling, the GSI did not change 
significantly between the early and late perinucleus 
stages (Table 3). After that, the GSI was significantly 
greater in each succeeding stage of maturity (Table 
3). Figure 5A shows that GSI remained low until fish 
reached lengths over 45 cm. GSI was significantly 
greater in mature males than in immature males 
