U.S. Department 
of Commerce 
Donald L. Evans 
Secretary 
National Oceanic 
and Atmospheric 
Administration 
Scott B. Gudes 
Acting Under Secretary for 
Oceans and Atmosphere 
National Marine 
Fisheries Service 
William T. Hogarth 
Acting Assistant Administrator 
for Fisheries 
Scientific Editor 
Dr. John V. Merriner 
Editorial Assistant 
Sarah Shoffler 
Center for Coastal Fisheries and Habitat Research, NOS 
101 Pivers Island Road 
Beaufort, NC 28516 
The Fishery Bulletin (ISSN 0090-0656) 
is published quarterly by the Scientific 
Publications Office, National Marine Fish- 
eries Service, NOAA, 7600 Sand Point Way 
NE, BIN C15700, Seattle, WA 98115-0070. 
Periodicals postage is paid at Seattle, WA, 
and at additional mailing offices. POST- 
MASTER: Send address changes for sub- 
scriptions to Fishery Bulletin, Superin- 
tendent of Documents, Attn.: Chief, Mail 
List Branch, Mail Stop SSOM, Washing- 
ton, DC 20402-9373. 
Although the contents of this publica- 
tion have not been copyrighted and may 
be reprinted entirely, reference to source 
is appreciated. 
The Secretary of Commerce has deter- 
mined that the publication of this peri- 
odical is necessary according to law for 
the transaction of public business of this 
Department. Use of funds for printing of 
this periodical has been approved by the 
Director of the Office of Management and 
Budget. 
For sale by the Superintendent of 
Documents, U.S. Government Printing 
Office, Washington, DC 20402. Subscrip- 
tion price per year: $45.00 domestic and 
$56.25 foreign. Cost per single issue: 
$28.00 domestic and $35.00 foreign. See 
back for order form. 
Managing Editor 
Sharyn Matriotti 
National Marine Fisheries Service 
Scientific Publications Office 
7600 Sand Point Way NE, BIN C 15700 
Seattle, Washington 98115-0070 
Editorial Committee 
Dr. 
Dr. 
Dr. 
Dr. 
Dr. 
Dr. 
Dr. 
Dr. 
Andrew E. Dizon 
Harlyn O. Halvorson 
Ronald W. Hardy 
Richard D. Methot 
Theodore W. Pietsch 
Joseph E. Powers 
Harald Rosenthal 
Fredric M. Serchuk 
National Marine Fisheries Service 
University of Massachusetts, Boston 
University of Idaho, Hagerman 
National Marine Fisheries Service 
University of Washington, Seattle 
National Marine Fisheries Service 
Universitat Kiel, Germany 
National Marine Fisheries Service 
Fishery Bulletin web site: fishbull.noaa.gov 
The Fishery Bulletin carries original research reports and technical notes on investigations in 
fishery science, engineering, and economics. It began as the Bulletin of the United States Fish 
Commission in 1881; it became the Bulletin of the Bureau of Fisheries in 1904 and the Fishery 
Bulletin of the Fish and Wildlife Service in 1941. Separates were issued as documents through 
volume 46; the last document was No. 1103. Beginning with volume 47 in 1931 and continuing 
through volume 62 in 1963, each separate appeared as a numbered bulletin. A new system 
began in 1963 with volume 63 in which papers are bound together in a single issue of the 
bulletin. Beginning with volume 70, number 1, January 1972, the Fishery Bulletin became a 
periodical, issued quarterly. In this form, it is available by subscription from the Superintendent 
of Documents, U.S. Government Printing Office, Washington, DC 20402. It is also available free in 
limited numbers to libraries, research institutions, State and Federal agencies, and in exchange 
for other scientific publications. 
