FISII PARASITES COLLECTED AT WOODS HOLE. 
299 
The following dimensions in millimeters were taken from alcoholic specimen: Diameter of head, 
lateral 1.95, marginal, 1.76; length of bothrial portion 1.8; distance from anterior end to where neck 
begins to -diminish 4; thickness of neck just back of bothria 1.4; distance to first distinct segments 
150; breadth of first distinct segments 0.84; length of last segments 0.56, breadth 0.9, thickness 0.37 ; 
length of head and nock 6. Length of large specimen in life, 250 mm. Small specimens not measured 
in life. The longest preserved small specimens are 18 mm. in length. A few measurements were 
made of the head of one in life, as follows: Breadth, bothria being extended nearly at right angles 
to axis 1.33; length of head proper, about 0.37; distance from anterior end to base of neck 1.33; 
diameter of neck just behind the bothria 0.46, diameter just before it begins to abruptly diminish 
0.36; breadth just back of neck 0.24 ; length of posterior segments 0.32, breadth 0.65. 
In the small specimens the first indication of segments, which appear as faint transverse aunu- 
lations, is about 8 mm. back of the head. The last segments are immature. In general proportions 
and shape they resemble the segments of the large specimen. 
The principal difference between the large specimen and the small ones is in the appearance of 
the head rather than in any essential dissimilarity of the bothria. In both the bothria are in pairs, 
and the pairs are on the sides of the head which correspond with the margins'of the body. In the 
alcoholic specimens the bothria are seen to be arranged in pairs, but the auriculate parts are directed 
in opposite directions, so that the two auriculate portions which are seen on the same side of the head 
really belong to different pairs of bothria (fig. 93). 
In large and small specimens alike the anterior part of a bothrium consists of a strong muscular 
sucker, shaped like a horseshoe, with the break in its border turned toward the posterior tip of the 
bothrium. The latter in the small specimens stands out as an auriculate appendage nearly at right 
angles to the axis of the body, while in the large specimens they are appressed. The neck in each 
case is thicker than the anterior part of the body, being, in fact, nearly cylindrical for a short dis- 
tance back of the head, where it diminishes in thickness, and, in the large specimen, also in breadth, 
rather abruptly. This cylindrical neck in the large specimen, proportionally to the head and body, 
is much larger than in the small specimens. The enlargement appears to affect the axial part of 
head also, thus filling in the interbothrial spaces and making the bothria sessile instead of prominent, 
as in the smaller ones. 
The genus Monorygma. is suggested by this species, and indeed Monticelli places the genus Calyp- 
trohothnnm near that genus. The head terminates abruptly without an eminence of any kind, which 
excludes the genus Monorygma. Again, the muscular auxiliary sucker on the front end of Iho bothria 
is of altogether different character from the auxiliary acetabulum of Phyllobothrium. 
Sections were made of several of the posterior segments of the large specimen, and, while the 
segments are immature, the general arrangement of the reproductive organs could be made out. The 
cirrus-pouch is pyriform and lies near one of the lateral margins, where it opens near middle of the 
length of proglottis. Within the bulb lie several convolutions of the vas deferens. The retracted 
cirrus was minute and not fully developed. A granular appearance on its walls suggested what might 
later develop into spines. The globular testicules occupy central portion of proglottis, mainly from 
a little behind the middle to anterior border. The Vagina opens in front of the cirrus in a common 
genital cloaca. The vitelline glands are voluminous and lie along the lateral margins. The ovary 
was identified as a smallish, lobulated mass of nuclei lying near the posterior margin of the proglottis, 
and staining somewhat differently from the vitelline glands. All the organs were for the most part 
masses of nuclei, staining deeply in carmine and presenting few differences. In the center of the 
segments was a mass of nuclei, some of which appeared to bo traveling to the vitellaria, and others 
forming the vas deferens and uterus. The latter, or what was so interpreted, appeared as a relatively 
large open space surrounded by a clustering mass of nuclei. 
Sections of posterior segments from the small specimens show testicules already begun and the 
rudiment of a cirrus-poucli. 
The neck, when sectioned, is seen to enlarge from the anterior part of the body by the expansion 
of the inner parenchyma, which consists of loosely intersecting libers with wide meshes, through 
which the longitudinal vessels pass in strong spirals. In the peripheral portions the longitudinal 
muscle fibers arc very strongly developed. Nuclei are sparse in the central portion of the neck except 
in the vicinity of the spiral longitudinal vessels. 
The most obvious difference between this species and Monticelli ’s species is in the character of 
the neck; in C. riggii the neck merges imperceptibly into the body, while in C. occidental a the neck is 
much thicker than the body and narrows rather abruptly a short distance back ot the head. 
