208 
Fishery Bulletin 95(2), 1997 
as the crew of the Ocean Hope III for their assistance 
and camaraderie that helped make this study pos- 
sible and enjoyable under adverse conditions. Data 
on sediment type, biomass, and abundance of inverte- 
brates were provided by H. M. Feder, Institute of Ma- 
rine Science, University of Alaska Fairbanks. We ap- 
preciate the assistance of R. Baxter and A. E. Peden in 
identifying some of the fishes. Finally, we thank A. V. 
Tfyler, S. C. Jewett, H. M. Feder, and unknown review- 
ers for comments that led to considerable improvements 
in the manuscript. This study was funded by the Alaska 
Outer Continental Shelf Region of the Minerals Man- 
agement Service, U. S. Department of the Interior, 
Anchorage, Alaska, Contract 14-35-0001-30559. 
Literature cited 
Alverson, D. L., and N. J. Wilimovsky. 
1966. Fishery investigations of the southeastern Chukchi 
Sea. In N. J. Wilimovsky and J. N. Wolfe (eds.). Environ- 
ment of the Cape Thompson region, Alaska, p. 843- 
860. U.S. Atomic Energy Commission, Washington, DC. 
Bianchi, G. 
1992a. Demersal assemblages of the continental shelf and 
upper slope of Angola. Mar. Ecol. Prog. Ser. 81:101-120. 
1992b. Study of the demersal assemblages of the continen- 
tal shelf and upper slope off Congo and Gabon Based on 
the trawl surveys of the RV ‘Dr. Fridtjof Nansen.’ Mar. 
Ecol. Prog. Ser. 85:9-23. 
Clifford, H. T., and W. Stevenson. 
1975. An introduction to numerical classification. Aca- 
demic Press, New York, NY, 229 p. 
Colvocoresses, J. A., and J. A. Musick. 
1984. Species associations and community composition of 
Middle Atlantic Bight continental shelf demersal 
fishes. Fish. Bull. 82(21:295-313. 
Day, D. S., and W. G. Pearcy. 
1968. Species associations of benthic fishes on the continen- 
tal shelf off Oregon. J. Fish. Res. Board Can. 25:2665-2675. 
Fargo, J., and A. V. Tyler. 
1991. Sustainability of flatfish-dominated fish assemblages 
in Hecate Strait, British Columbia, Canada. Neth. J. Sea 
Res. 27 (3/41:237-253. 
Feder, H. M., N. R. Foster, S. C. Jewett, T. J. Weingartner, 
and R. Baxter. 
1994. Molluscs in the northeastern Chukchi Sea. Arctic 
47:145-163. 
Frost, K. J., and L. F. Lowry. 
1983. Demersal fishes and invertebrates trawled in the north- 
eastern Chukchi and western Beaufort seas 1976-1977. 
U.S. Dep. Commer., NOAATfcch. Rep. NMFS-SSRF-764, 22 p. 
Jahn, A. E., and R. H. Backus. 
1976. On the mesopelagic fish faunas of Slope Water, Gulf 
Stream and northern Sargasso Sea. Deep-Sea Res. 
23:223-234. 
Johnson, W. R. 
1 989. Current response to wind in the Chukchi Sea: a regional 
coastal upwelling event. J. Geophys. Res. 94:2057-2064. 
Lance, G. N., and W. T. Williams. 
1967. A general theory of classificatory sorting strategies. 
1: Hierarchical systems. Comput. J. 9:373380. 
Mahon, R., and R. W. Smith. 
1989. Demersal fish assemblages on the Scotian Shelf, 
Northwest Atlantic: spatial distribution and persistence. 
Can. J. Fish. Aquat. Sci. 46 (suppl. 1:134-152. 
McKelvie, D. S. 
1985. Discreteness of pelagic faunal regions. Mar. Biol. 
88:125-133. 
Naidu, A. S. 
1988. Marine surficial sediments. Section 1.2 in Bering, 
Chukchi, and Beaufort seas: coastal and ocean zones stra- 
tegic assessment data atlas, p. 1.4. U.S. Dep. Commer., 
NOAA, Strategic Assessment Branch, Ocean Assessment 
Division, Rockville, MD. 
Overholtz, W. J., and A. V. Tyler. 
1985. Long-term responses of the demersal fish assem- 
blages of Georges Bank. Fish. Bull. 83(4):507-520. 
Pearcy, W. G. 
1978. Distribution and abundance of small flatfishes and 
other demersal fishes in a region of diverse sediments and 
bathymetry off Oregon. Fish. Bull. 76(31:629-640. 
Pielou, E. C. 
1977. Mathematical ecology. John Wiley and Sons, New 
York, NY, 385 p. 
Pruter, A. T., and D. L. Alverson. 
1962. Abundance, distribution and growth of flounders in 
the southeastern Chukchi Sea. J. Cons. Cons. Int. Explor. 
Mer 27:81-99. 
Scott, J. S. 
1982. Selection of bottom type by groundfishes of the 
Scotian Shelf. Can. J. Fish. Aquat. Sci. 39:943-947. 
Sharma, G. D. 
1979. The Alaskan shelf: hydrographic, sedimentary and 
geochemical environment. Springer- Verlag, New York, 
NY, 498 p. 
Siegel, S., and N. J. Castellan Jr. 
1988. Nonparametric statistics for the behavioral 
sciences. McGraw-Hill Book Co., New York, NY, 399 p. 
Smith, R. L., W. E. Barber, M. Vallarino, E. Barbour, and 
E. Fitzpatrick. 
In press, a. Biology of the Bering flounder, Hippoglossoides 
robustus , from the northeastern Chukchi Sea. In J. B. 
Reynolds (ed.l. Fish ecology ofArctic North America. Am. 
Fish. Soc. Symp. 
Smith, R. L., W. E. Barber, M. Vallarino, J. A. Gillispie, 
and A. Ritchie. 
In press, b. Biology of Gymnocanthus tricuspis, the Arctic 
staghorn sculpin, from the northeastern Chukchi Sea. In 
J. B. Reynolds (ed.), Fish ecology of Arctic North 
America. Am. Fish. Soc. Symp. 
Wakabayashi, K., R. G. Bakkala, and M. S. Alton. 
1985. II: Methods of the U.S. -Japan demersal trawl 
surveys. In R. G. Bakkala and K. Wakabayashi (eds.), 
Results of cooperative U.S. -Japan groundfish investiga- 
tions in the Bering Sea during May-August 1979, p. 7— 
29. Int. North Pac. Fish. Comm. Bull. 44. 
Weinberg, K. L. 
1994. Rockfish assemblages of the middle shelf and upper 
slope off Oregon and Washington. Fish. Bull. 92(3 ):620-632. 
Weingartner, T. J. 
In press. The physical oceanography of the northeastern 
Chukchi Sea: a review. In J. B. Reynolds (ed.), Fish ecol- 
ogy ofArctic North America. Am. Fish. Soc. Symp. 
Zar, J. H. 
1984. Biostatistical analysis. Prentice Hall, Englewood 
Cliffs, NJ, 718 p. 
