BIOLOGICAL STUDY OF CHESAPEAKE BAY WATERS 
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CONFINED ALMOST TO INNER BAY 
[North of Maryland and Virginia linej 
*Canuella elongata Wilson, *Cletodes longicaudalus (Boeck), * Dactylopusia brevicornis (Claus), 
Ectinosoma normani T. and A. Scott, Harpacticus liltoralis G. O. Sars, *Robertsonia chesapcakensis 
Wilson, *Tachidius littoralis Poppe, Eurytemora americana Williams, E. hirundoides (Nordquist), 
and *Mesocyclops gracilis (Lilljeborg). Three other species, *Hemicyclops americanus Wilson, 
Candacia armata Boeck, and *Bomolochus eminens Wilson occurred in very small numbers at one or 
two areas. 
CONFINED ALMOST TO OUTER BAY 
[South of Maryland and Virginia line] 
Alleutha depressa Baird, Calanus finmarchicus (Gunnerus), Corycaeus carinatus Giesbrecht, C. 
elongatus Claus, *C. venustus Dana, *Cryptopontius gracilis Wilson, *Oithona spiniroslris Claus, 
Oncaea minula Giesbrecht, *Labidocera wollastoni (Lubbock), Pontella meadii Wheeler, Temora 
longicornis (Miiller), *T. turbinata (Dana), Tisbe furcata (Baird), Microsetella norvegica (Boeck), 
Harpacticus chelifer (0. F. Midler), and Diosaccus tenuicornis (Claus). In addition the following 
species, in very small numbers, were found at one or two areas: * Temora discaudata Giesbrecht, 
Pontella pennata Wilson, and Caligus schistonyx Wilson. 
FOUND ONLY OUTSIDE OF THE BAY 
[Between the capes and the 100-fathom line] 
* Amallophora brevicornis G. O. Sars, Anomalocera patersoni Templeton, *Calanus helgolandicus 
(Claus), Centropages bradyi Wheeler, Euchaeta norvegica Boeck, Mecynocera clausii I. C. Thompson, 
Metridia lucens Boeck, *Pontella atlantica (Milne Edwards), Rhincalanus nasutus Giesbrecht, 
Clytemnestra rostrata (Brady), Macrosetella gracilis (Dana), *Corycaeus lubbockii Giesbrecht, *C. 
speciosus Dana, Oithona plumifera Baird, Oncaea venusta Philippi, Sapphirina gemma Dana, and *S. 
sinuicauda Brady, Corycaeus robustus Giesbrecht, C. rostratus Claus. 
These lists total 64 species: 13 universal, or almost so, over the bay; 13 almost 
confined to the inner bay; 19 almost confined to the outer bay; and 19 outside of the 
bay only. Not less than 19 of the 26 species listed under "universal ” and "inner bay ” 
occur in such bodies of water as Chesapeake Bay or at least frequent coastal waters 
in other parts of the world, and these 19 do not include the new species and the 
species whose distribution is very little known. Of those listed under "outer bay” 
(19) a much smaller number (6) are characteristic of estuaries in other regions, while 
those listed for "outside of the bay” (19) include not more than 3 or 4 species which 
are recorded as being estuarine forms. In other words, the number of estuarine 
forms found during our cruises decreases, passing from the inner bay to the region 
outside of the capes. On the whole, from this point of view, the distribution of the 
free-swimming copepods found by us in Chesapeake Bay and the region immediately 
outside of the bay is much like that of the same copepods in other parts of the world. 
Twenty-two of the species listed above (those with an asterisk) are either new 
species (named and described by Wilson but still unpublished) or those which have 
not been reported hitherto from the eastern coast of North America. 
There are included in the complete list 1 species, Bomolochus eminens, which is 
known to occur parasitically in the gill cavity of the false Spanish sardine, Clupanodon 
pseudohispanicus ; 1 species, Sapphirina gemma, caught free-swimming but known to 
be a commensal in Salpa; 1 species, S. sinuicauda, also caught free-swimming but 
probably a commensal in Salpa; 1 species, Caligus shistonyx, an external parasite on 
the menhaden, Brevoortia tyrannus; and 1 species, Mesocyclops gracilis which is a 
fresh -water form. 
Most of the copepods caught in Chesapeake Bay have been present in such small 
numbers that any extended discussion of their ecology is not permissible; but one 
