Misa et a!.: Establishing species-habitat associations for 4 eteline snappers 
299 
LLJ 
CO 
+ 
15 
12 
9 - 
6 - 
3 
0 
130 
25 
20 
15 
10 
5 
0 
i hard-high a hard-low □ soft-high □ soft-low ®BRFA 
Opakapaka (90-210 m) 
30 7 24 2 63 8 12 5 6 31 25 21 25 7 78 9 15 14 5 43 0 25 0 8 33 8 14 13 7 42 80 94 81 35 
Kalekale (180-270 m) 
22 17 14 10 63 5 14 7 3 29 19 10 15 7 51 31 12 18 12 73 0 77 0 5 82 16 0 7 1 24 93 130 61 38 
Habitat 
Mean relative abundance (MaxNo) with standard error (SE) by location of bottomfish restricted 
fishing area (BRFA; i.e., combined area inside and outside a BRFA), by habitat type combined 
for all BRFA locations (habitat), and by habitat type in each BRFA location (BRFAxhabitat) for 
Opakapaka (Pristipomoides filamentosus), Kalekale (P. sieboldii), Onaga (Etelis coruscans), and 
Ehu (E. carbunculus) within the preferred depths of each species in the main Hawaiian Islands. 
A baited stereo-video camera system (BotCam) was used to collect data from May 2007 to June 
2009. Columns with the same letter (uppercase type for BRFA; lowercase, bold, italic type for hab- 
itat; lowercase type for BFRAxhabitat) are not significantly different from each other (P>0.05, 
post hoc permutational analysis of variance [PERMANOVA]). The number below each column is 
the number of BotCam deployments. The 4 habitat classifications used in our study were derived 
from data collected with multitaeam sonar systems: hard substrate with high slope (hard-high), 
hard substrate with low slope (hard-low), soft substrate with high slope (soft-high), and soft sub- 
strate with low slope (soft-low). The 8 sampled BRFAs were (B) Niihau, (D) Kaena, (E) Makapuu, 
(F) Penguin Bank, (H) Pailolo Channel, and (L) Hilo. Error bars indicate ±1 SE of the mean. 
