Frable et al: A new species of Sebastes 
375 
feedback and support, our undergraduate helpers, A. 
Martin and E. Peterson, A. Hollmann (UW) for eluci- 
dating Ancient Greek etymology and for assisting with 
naming the new species, and the Oregon Coast Aquari- 
um for access to exhibits. Support was provided by Na- 
tional Science Foundation Division of Biological Infra- 
structure NSF DBI 1057452 and U.S. Fish and Wildlife 
Service Federal Assistance Award OR F-186-R-9. 
Literature cited 
An, H. S., J. Y. Park, M.-J. Kim, E. Y. Lee, and K. K. Kim. 
2009. Isolation and characterization of microsatellite 
markers for the heavily exploited rockfish Sebastes 
schlegeli, and cross-species amplification in four related 
Sebastes spp. Conserv. Genet. 10:1969-1972. 
Ayres, W. 0. 
1854. Description of new fishes from California. Proc. 
Calif. Acad. Nat. Sci. 1:3-22. 
1862. Descriptions of fishes believed to be new. Proc. Ca- 
lif. Acad. Nat. Sci. 2:206-218. 
Bean, T. H. 
1882. Notes on a collection of fishes made by Captain 
Henry E. Nichols, U.S.N., in British Columbia and 
southern Alaska, with descriptions of new species and a 
new genus (Delolepis). Proc. U.S. Natl. Mus. 4:463-474. 
Bohlke, E. B. 
1984. Catalog of type specimens in the ichthyological 
collection of the Academy of Natural Sciences of Phila- 
delphia. Special Publication 14, 216 p. Acad. Nat. Sci. 
Phil., PA. 
Burford, M. O. 
2009. Demographic history, geographical distribution 
and reproductive isolation of distinct lineages of blue 
rockfish (Sebastes mystinus), a marine fish with a high 
dispersal potential. J. Evol. Biol. 22:1471-1486. 
Burford, M. O., and G. Bernardi. 
2008. Incipient speciation within a subgenus of rockfish 
( Sebastosomus ) provides evidence of recent radiations 
within an ancient species flock. Mar. Biol. 154:701-717. 
Burford, M. O., G. Bernardi, and M. H. Carr. 
2011a. Analysis of individual year-classes of a marine 
fish reveals little evidence of first generation hybrids 
between cryptic species in sympatric regions. Mar. 
Biol. 158:1815-1827. 
Burford, M. O., M. H. Carr, and G. Bernardi. 
2011b. Age-structured genetic analysis reveals temporal 
and geographic variation within and between two cryp- 
tic rockfish species. Mar. Ecol. Prog. Ser. 442:201-215. 
Burford, M. O., and R. J. Larson. 
2007. Genetic heterogeneity in a single year-class from 
a panmictic population of adult blue rockfish (Sebastes 
mystinus). Mar. Biol. 151:451-465. 
Burnaby, T. P. 
1966. Growth-invariant discriminant functions and gen- 
eralized distances. Biometrics 22:96-110. 
Clemens, W. A., and G. V. Wilby. 
1961. Fishes of the Pacific coast of Canada, 2 nd ed. Fish. 
Res. Board Can. Bull. 68, 443 p. 
Cope, J. M. 
2004. Population genetics and phylogeography of the 
blue rockfish (Sebastes mystinus ) from Washington to 
California. Can. J. Fish. Aquat. Sci. 61:332-342. 
Crow, K. D., H. Munehara, and G. Bernardi 
2010. Sympatric speciation in a genus of marine fish- 
es. Mol. Ecol. 19:2089-2105. 
Earl, D. A., and B. M. vonHoldt. 
2012. STRUCTURE HARVESTER: a website and pro- 
gram for visualizing STRUCTURE output and imple- 
menting the Evanno method. Conserv. Genet. Res. 
4:359-361. 
Eigenmann, C. H., and C. H. Beeson. 
1893. Preliminary note on the relationship of the spe- 
cies usually united under the generic name Sebas- 
todes. Am. Nat. 27:668-671. [Available at website.] 
Eschmeyer, W. N., E. S. Herald, and H. Hammann. 
1983. A field guide to Pacific Coast fishes of North Amer- 
ica, 336 p. Houghton Mifflin Co., Boston, MA. 
Falush, D., M. Stephens, and J. K. Pritchard. 
2003. Inference of population structure using multilocus 
genotype data: linked loci and correlated allele frequen- 
cies. Genetics 164:1567-1587. 
Gharrett, A. J., A. P. Matala, E. L. Peterson, A. K. Gray, Z. Li, 
and J. Heifetz. 
2005. Two genetically distinct forms of Rougheye rock- 
fish (Sebastes aleutianus ) are different species. Trans. 
Am. Fish. Soc. 134:242-260. 
Girard, C. F. 
1856. Contributions to the ichthyology of the western 
coast of the United States, from specimens in the mu- 
seum of the Smithsonian Institution. Proc. Acad. Nat. 
Sci. Phila. 8:131-137. 
Gomez-Uchida, D., E. A. Hoffman, W. R. Ardren, and M. A. Banks. 
2003. Microsatellite markers for the heav- 
ily exploited canary (Sebastes pinniger) and oth- 
er rockfish species. Mol. Ecol. Notes 3:387-389. 
Goode, G. B. 
1884. The fisheries and fishery industries of the United 
States. Section I: natural history of useful aquatic ani- 
mals, 895 p. Government Printing Office, Washington, 
D.C. 
Hammer, 0., D. A. T. Harper, and P. D. Ryan. 
2001. PAST: paleontological statistics software package 
for education and data analysis. Palaeontologia Elec- 
tronica 4(1): 1—9. [Available at website.] 
Hart, J. L. 
1973. Pacific fishes of Canada. Fish. Res. Board Can. Bull. 
180, 740 p. 
Helvey, M. 
1982. First observations of courtship behavior in rock- 
fish, genus Sebastes. Copeia 1982:763-770. 
Hyde, J. R., and R. D. Vetter. 
2007. The origin, evolution, and diversification of rock- 
fishes of the genus Sebastes (Cuvier). Mol. Phylogen. 
Evol. 44:790-811. 
Hyde, J. R., C. A. Kimbrell, J. E. Budrick, E. A. Lynn, and R. 
D. Vetter. 
2008. Cryptic speciation in the vermilion rockfish ( Se- 
bastes miniatus) and the role of bathymetry in the spe- 
ciation process. Mol. Ecol. 17:1122-1136. 
Jordan, D. S., and B. W. Evermann. 
1898. The fishes of North and Middle America: a de- 
scriptive catalogue of the species of fish-like verte- 
brates found in the waters of North America, north of 
the Isthmus of Panama, part II. Bull. U.S. Natl. Mus. 
47:1241-2183. 
1900. The fishes of North and Middle America: a de- 
scriptive catalogue of the species of fish-like verte- 
