Martinho et al.: Comparative feeding ecology of two elasmobranch species, Squaius blainville and Scyliorhinus canicula 
83 
Avsar, D. 
2001. Age, growth, reproduction and feeding of the 
spurdog ( Squaius acanthias Linnaeus, 1758) in the 
South-eastern Black Sea. Estuar. Coast. Shelf Sci. 
52:269-278. 
Baeta, F., M. Batista, A. Maia, M. J. Costa, and H. N. Cabral. 
2010. Elasmobranch bycatch in a trammel net fish- 
ery in the Portuguese west coast. Fish. Res. 102: 
123-129. 
Bowen, W. D. 
1997. Role of marine mammals in aquatic ecosys- 
tems. Mar. Ecol. Prog. Ser. 158:267-274. 
Braccini, J. M., B. M. Gillanders, and T. 1. Walker. 
2005. Sources of variation in the feeding ecology of 
the piked spurdog ( Squaius megalops)', implications for 
inferring predator-prey interactions from overall dietary 
composition. ICES J. Mar. Sci. 62:1076-1094. 
Cannizzaro, L. P. Rizzo, D. Levi, and S. Gancitano. 
1995. Age-determination and growth of Squaius blainvil- 
lei (Risso, 1826). Fish. Res. 23:113-125. 
Capape, C. 
1974. Contribution a la biologie des Scyliorhinidae des 
cotes Tunisiennes. Annales de L’Institut Michel Pacha 
7: 13-29. [In French.] 
1975. Etude du regime alimentaire de Squaius bainvil- 
lei (Risso, 1826) des cotes Tunisiennes. Bull. Inst. 
Natl. Sci. Tech. Oceanogr. Peche Salammbo 4:61-73. 
[In French.] 
1977. Contribution a la biologie des Scyliorhinidae des 
cotes Tunisiennes Scyliorhinus canicula , fecondite. repar- 
tition geographique et bathymetrique, sexualite, repro- 
duction. Bull. Office National des Peches de Tunisie 
(L. 1978) 1:83-101. [In French.] 
Capape, C., C. Reynaud, Y. Vergne, and J.-P. Quinard. 
2008. Biological observations on the smallspotted cat- 
shark Scyliorhinus canicula (Chondrichthyes: Scyliorh- 
inidae) off the Languedocian coast (southern France, 
northern Mediterranean). Pan-Am. J. Aquat. Sci. 
3:282-289. 
Carbonell, A., F. Alemany, P. Merella, A. Quetglas, and E. 
Roman. 
2003. The by-catch of sharks in the western Mediter- 
ranean (Balearic Islands) trawl fishery. Fish. Res. 
61:7-18. 
Christensen, V., and D. Pauly. 
1992. ECOPATH II — a software for balancing steady- 
state ecosystem models and calculating network char- 
acteristics. Ecol. Model. 61:169-185. 
Coelho, R., K. Erzini, L. Bentes, C. Correira, P. G Lino, P. 
Monteiro, J. Ribeiro, and J. M. S. Gongalves. 
2005. Semi-pelagic longline and trammel net elasmo- 
branch catches in southern Portugal: catch composition, 
catch rates and discards. J. Northwest Atl. Fish. Sci. 
35:531-537. 
Colwell, R. K. 
2009. EstimateS: Statistical estimation of species rich- 
ness and shared species from samples, V.8.2. User’s 
guide and application. [Available from http://purl.oclc. 
org/estimates, accessed July 2011.] 
Cortes, E. 
1999. Standardized diet compositions and trophic levels 
of sharks. ICES J. Mar. Sci. 56:707-717. 
Craik, J. C. A. 
1978. An annual cycle of vitellogenesis in the elasmo- 
branch Scyliorhinus canicula. J. Mar. Biol. Assoc. U.K. 
58:719-726. 
Crothers, J. 
1983. A key to the crabs and crab-like animals of British 
inshore waters. Field Stud. 5:753-806. 
Demirhan, S. A., and K. Seyhan. 
2007. Life history of spiny dogfish, Squaius acanthias 
(L. 1758), in the southern Black Sea. Fish. Res. 85: 
210-216. 
Domi, N., J. M. Bouquegneau, and K. Das. 
2005. Feeding ecology of five commercial shark species 
of the Celtic Sea through stable isotope and trace metal 
analysis. Mar. Environ. Res. 60:551-569. 
Ellis, J. R„ M. G. Pawson, and S. E. Shackley. 
1996. The comparative feeding ecology of six species 
of shark and four species of ray (Elasmobranchii) in 
the North-East Atlantic. J. Mar. Biol. Assoc. U.K. 
76:89-106. 
Fauvel, P. 
1923. Polyehetes Errantes. In Faune de France, vol. 5 
(P. Lechevalier, ed.), 488 p. Federation Frangaise des 
Societes de Sciences Naturelles, Paris. [In French.] 
1927. Polyehetes Sedentaires. In Faune de France, 
vol. 16 (P. Lechevalier, ed.), 494 p. Federation Fran- 
gaise des Societes de Sciences Naturelles, Paris. 
[In French.] 
Ferry, L. A., and G. M. Cailliet. 
1996. Sample size and data analysis: are we charac- 
terizing and comparing diet properly? In GUTSHOP 
’96: feeding ecology and nutrition in fish: symposium 
proceedings (D. MacKinley and K. Shearer, eds.), p. 
71-80. Am. Fish. Soc. Physiol. Sect, and Int. Congress 
Biol. Fishes, July 1996, San Francisco, CA. 
Filiz, H., and E. Taskavak. 
2006. Sexual dimorphism in the head, mouth, and body 
morphology of the smallspotted catshark, Scyliorhinus 
canicula (Linnaeus, 1758) (Chondrichthyes: Scyliorhini- 
dae) from Turkey. Acta Adriat. 47:37-47. 
Froese, R., and D. Pauly (eds). 
2010. FishBase. www.fishbase.org, accessed December 
2010 . 
Hayward, P. J., and J. S. Rylancl. 
1995. Handbook of the marine fauna of North-West 
Europe, 800 p. Oxford Univ. Press, Oxford. 
Herndon, A., V. F. Gallucci, D. Demaster, and W. Burke. 
2010. The case for an international commission for the 
conservation and management of sharks (ICCMS). Mar. 
Policy 34:1239-1248. 
Hyslop, E. J. 
1980. Stomach contents analysis: a review of methods 
and their application. J. Fish Biol. 17:415-429. 
Juanes, F., J. A. Buckel, and F. S. Scharf. 
2001. Predatory behaviour and selectivity of a primary 
piscivore: comparison of fish and non-fish prey. Mar. 
Ecol. Prog. Ser. 217:157-165. 
Kousteni, V., and P. Megalofonou. 
2011. Reproductive biology and embryonic development 
of Squaius blainvillei in the eastern Mediterranean 
Sea. Sci. Mar. 75:237-249. 
Kimber, J. A., D. W. Sims, P. H. Bellamy, and A. B. Gill. 
2009. Male-female interactions affect foraging behaviour 
within groups of small-spotted catshark, Scyliorhinus 
canicula. Anim. Behav. 77:1435-1440. 
Linton, R. L., R. W. Davies, and F. J. Wrona. 
1981. Resource utilization indices; an assessment. J. 
Anim. Ecol. 50:283-293. 
Olaso, I., F. Velasco, and N. Perez. 
1998. Importance of discarded blue whiting (Microme- 
