160 
Fishery Bulletin 111(2) 
Holts, D. B., and D. W. Bedford. 
1993. Horizontal and vertical movements of the shortfin 
mako shark, Isurus oxyrinchus, in the southern Califor- 
nia bight. Aust. J. Mar. Freshw. Res. 44:901-909. 
Joung, S. J., and H. H Hsu. 
2005. Reproduction and embryonic development of the 
shortfin mako, Isurus oxyrinchus Rafinesque, 1810, in 
the northwestern Pacific. Zool. Stud. 44:487-496. 
Kusher, D. I., S. E. Smith, and G. M. Cailliet. 
1992. Validated age and growth of the leopard shark, 
Triakis semifasciata, with comments on reproduc- 
tion. Environ. Biol. Fishes 35:187-203. 
MacDonald, P. D. M., and T. J. Pitcher. 
1979. Age-groups from size-frequency data: a versatile 
efficient method of analyzing distribution mixtures. J. 
Fish. Res. Board Can. 36:987-1001. 
Maia, A., N. Queiroz, H. N. Cabral, A. M. Santos, and J. P. 
Correia. 
2007. Reproductive biology and population dynamics of 
the shortfin mako, Isurus oxyrinchus Rafinesque, 1810, 
off the southwest Portuguese coast, eastern North At- 
lantic. J. Appl. Ichthyol. 23:246-251. 
Mollet, H. F., G. Cliff, H. L. Pratt, and J. D. Stevens. 
2000. Reproductive biology of the female shortfin mako, 
Isurus oxyrinchus Rafinesque, 1810, with comments on 
the embryonic development of lamnoids. Fish. Bull. 
98:299-318. 
Monaghan, J. P. 
1993. Comparison of calcein and tetracycline as chemi- 
cal markers in summer flounder. Trans. Am. Fish. Soc. 
122:298-301. 
Murphy, M. D., D. H. Adams, D. M. Tremain, and B. L. Winner. 
1998. Direct validation of ages determined for adult 
black drum, Pogonias cromis , in east-central Flori- 
da, with notes on black drum migration. Fish. Bull. 
96:382-387. 
Natanson, L. -J., N. E. Kohler, D. Ardizzone, G. M. Cailliet, S. 
P Wintner, and H. F. Mollet. 
2006. Validated age and growth estimates for the short- 
fin mako, Isurus oxyrinchus , in the North Atlantic 
Ocean. Environ. Biol. Fishes 77:367-383. 
Natanson, L. J., S. P. Wintner, F. Johansson, A. Piercy, P. 
Campbell, A. D Maddalena, S. J. B. Gulak, B. Human, F. C. 
Fulgosi, D. A. Ebert, F. Hemida, F. H. Mollen, S. Vanni, G. 
H. Burgess, L. J. V. Compagno, and A. Wedderburn-Maxwell. 
2008. Ontogenetic vertebral growth patterns in the 
basking shark Cetorhinus maximus. Mar. Ecol. Prog. 
Ser. 361:267-278. 
Okamura, H., and Y. Semba. 
2009. A novel statistical method for validating the peri- 
odicity of vertebral growth band formation in elasmo- 
branch fishes. Can. J. Fish. Aquat. Sci. 66:771-780. 
PFMC (Pacific Fishery Management Council). 
2011. Fishery management plan for U.S. West Coast 
fisheries for highly migratory species. As amended 
through Amendment 2, 106 p. Pacific Fishery Man- 
agement Council, Portland, OR. [Available from 
http://www.pcouncil.org/highly- migratory -species/ 
fishery-management-plan-and-amendments/.] 
Parker, H. W., and F. C. Stott. 
1965. Age, size and vertebral calcification in the basking 
shark, Cetorhinus maximus (Gunnerus). Zool. Meded. 
40:305-319. 
Pauly, D. 
2002. Growth and mortality of the basking shark Ceto- 
rhinus maximus and their implications for management 
of whale sharks Rhincodon typus. In Elasmobranch 
biodiversity, conservation and management. Proceed- 
ings of the international seminar and workshop; Sabah, 
Malaysia, July 1997 (S. L. Fowler, T. Reid, and F. A. Dip- 
per, eds.), p. 199-208. Occasional Papers of the IUCN 
Survival Commission, no. 25. Int. Union Conserv. Na- 
ture, Gland, Switzerland. 
Pratt, H. L., Jr., and J. G. Casey. 
1983. Age and growth of shortfin mako, Isurus oxyrin- 
chus i, using four methods. Can. J. Fish. Aquat. Sci. 
40:1944-1957. 
R Development Core Team. 
2008. R: a language and environment for statistical 
computing. R Foundation for Statistical Computing. 
[Available from: http://R-project.org, accessed November 
2008.] 
Ribot-Carballal, M. C., F. Galvan-Magana, and C. Quinonez- 
Velazquez. 
2005. Age and growth of the shortfin mako shark, Isurus 
oxyrinchus , from the western coast of Baja California 
Sur, Mexico. Fish. Res. 75:14-21. 
Semba, Y., H. Nakano, and I. Aoki. 
2009. Age and growth analysis of the shortfin mako 
shark, Isurus oxyrinchus , in the western and central 
North Pacific Ocean. Environ. Biol. Fishes 84:377-391. 
Semba, Y. I. Aoki, and K. Yokawa. 
2011. Size at maturity and reproductive traits of short- 
fin mako, Isurus oxyrinchus , in the western and central 
North Pacific. Mar. Freshw. Res. 62:20-29. 
Smith, S. E., D. W. Au, and C. Show. 
1998. Intrinsic rebound potentials of 26 species of Pacific 
sharks. Mar. Freshw. Res. 49:663-678. 
Smith, S. E., R. A. Mitchell, and D. Fuller. 
2003. Age-validation of a leopard shark recaptured after 
20 years. Fish. Bull. 101:194-198. 
Stevens, J. D. 
2008. The biology and ecology of the shortfin mako 
shark, Isurus oxyrinchus. In Sharks of the open ocean: 
biology, fisheries and conservation (M. D. Camhi, E. K. 
Pikitch, and E. A. Babcock, eds), p. 87-94. Blackwell 
Publ., Ltd., Oxford. 
Tanaka, S. 
1990. Age and growth studies on the calcified struc- 
tures of newborn sharks in the laboratory aquaria us- 
ing tetracycline. In Elasmobranchs as living resources: 
advances in the biology, ecology, systematics, and the 
status of the fisheries (H. L. Pratt Jr., S. H. Gruber, and 
T. Taniuchi, eds.), p. 189-202. NOAA Tech. Rep. NMFS 
90. 
Taylor, V. B., and D. W. Bedford. 
2001. Shortfin mako shark. In California living marine 
resources: a status report (W. S. Leet, C. M. DeWees, R. 
Klingbeil, and E. J. Larson, eds), p. 336-337. Publica- 
tion SG01-11, California Dep. Fish Game, Sacramento, 
CA. 
Winker, H., O. L. F. Weyl, A. J. Booth, and B. R. Ellender. 
2010. Validating and corroborating the deposition of two 
annual growth zones in asteriscus otoliths of common 
carp Cyprinus carpio from South Africa’s largest im- 
poundment. J. Fish Biol. 77:2210-2228. 
