Capossela et al.: Migratory and within-estuary behaviors of adult Paralichthys dentatus of the southern mid-Atlantic Bight 
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Figure 1 
(A) The distribution of acoustic receivers and water-quality sondes installed in the Wachapreague lagoon system for this study of 
Summer Flounder ( Paralichthys dentatus) behaviors between June 2007 and July 2008. The location of the Wachapreague lagoon 
system in the southern mid-Atlantic Bight is shown by the square in the smaller map. Regions specified in the text are Wacha- 
preague Inlet, upper channels, lower channels, and tidal flat. Receivers S3, S5-S9, Sll, and S12 provided supplemental data on 
the activity in regions outside of our study area. (B) In this map of acoustic receivers, each circle represents the approximate 
detection range (radius=35Q m) of the receivers deployed in the Wachapreague lagoon system. 
temperatures (Capossela et al., 2012). Photoperiod (i.e., 
day length) was acquired from tide prediction software 
(Jtides, vers. 4.9; http://www.arachnoid.com/JTides). 
Telemetry and tagging 
On 22 May 2007, 50 Summer Flounder (261—558 mm 
total length [TL] ) were captured at the study area 
by hook and line, identified from Murdy et al (1997), 
and immediately anesthetized with 60 mg L _1 AQUI- 
S (AQUI-S New Zealand Ltd., Lower Hutt, New Zea- 
land) to allow surgical implantation of individually 
coded 69-kHz transmitters (V9-2L-R64K; VEMCO Di- 
vision, AMIRIX Systems, Inc., Bedford, Canada) by 
using established procedures (Fabrizio and Pessutti, 
2007). Transmitters were 30 mm long and 9 mm in 
diameter and had a delay time of 60-180 s and a pro- 
jected 14-month battery life. All fish were tagged and 
released in the upper channels with the exception of 
a single fish that was captured, tagged, and released 
on the tidal flat. Before release, all fish were allowed 
to fully recover in an onboard aquarium that accom- 
modated total length, and externally tagged with an 
individually numbered T-bar anchor tag inserted near 
the caudal peduncle to alert anglers to report recap- 
tures. We considered all fish to be adults because Sum- 
mer Flounder can reach maturity at 240-300 mm TL 
(Morse, 1981). 
Summer Flounder migratory and within-estuary be- 
haviors were examined from 8 June 2007 until the last 
fish departed on 17 January 2008. We chose the start 
date (8 June 2007), which was approximately 2 weeks 
after the release of tagged fish, to limit the influence 
of any atypical activity patterns due to recovery from 
capture and surgery (Knights and Lasse, 1996; Rogers 
and White, 2007). We recorded fish locations with 31 
