Orr et al.: Systematics of North Pacific sand lances of the genu s Ammodytes and description of a new species 
155 
(70-105 mm), Gulf of Alaska, Uyak Bay, Kodiak Island, 
57.6°N, 154°W, 10 July 1941, A. C. DeLacy et al.; UW 
15610, 3 (90-123 mm), Alexander Archipelago, Shelikof 
Bay, Kruzof Island, 57.15°N, 135.8°W, 1 July 1960, Dry- 
foos; UW 16464, 4 (145-152 mm), 57.4167°N, 150.75°W, 
4 June 1958; UW 20642, 8 (72-85 mm), Kodiak Island, 
Ugak Bay, 57.4517°N, 152.7019°W, U2-301-117, 20 June 
1976, C. Harris and M. Hunter; UW 243, 1 (75 mm), 
Kodiak Island, Alitak, 56.9°N, 154.16°W, Crawford; UW 
26825, 1 (95 mm), Kodiak Island, Ugak Bay, 57.4517°N, 
152.701°W, 4 August 1923; UW 4002, 3 (70-90 mm), 
Kodiak Island, Lazy Bay, 56.9°N, 154.26°W, 8 May 
1932; UW 4202, 4 (85-90 mm), Wide Bay, 57.4°N, 
156.46°W, 31 May 1932; UW 45961, 1 (91 mm), 
57.515°N, 151.5496°W, 147 m depth, FV Vesteraalen, 
cruise 200101, haul 177, 28 June 2001, J. W. Orr; UW 
5301, 8 (112.0-120.2 mm), Prince of Wales Island, 
Tranquil Point in the Alexander Archipelago, 55°N, 
133°W, 22 July 1939, Franett. British Columbia: UW 
1734, 1 (83 mm), Hecate Strait, middle of Hecate Strait 
between Gander Island and Danger Rocks, 52.5°N, 
129. 5°W, 1 September 1931. Washington: UW 11329, 8 
(66.1-107.1 mm), Puget Sound, Meadow Point, Seattle, 
47.6°N, 122. 3°W, 29 April 1955, A. DeLacy and class; 
UW 116044-116054, 11 (61.2-87.1 mm), Puget Sound, 
48.7042°N, 123.014°W, 60-80 m depth, RV Centennial, 
site 10, sample V5, 6 November 2006, D. Gunderson 
and J. Blaine; UW 116055-116064, 10 (65.6-75.2 mm), 
Puget Sound, 48.69°N, 123.0161°W, 60-80 m depth, RV 
Centennial, site 28, sample V18, 6 November 2006, D. 
Gunderson and J. Blaine; UW 116065-116075, 11 
(65.0-75.2 mm), Puget Sound, 48.5325°N, 123.0214°W, 
60-80 m depth, RV Centennial, site 26, sample V16, 6 
November 2006, D. Gunderson and J. Blaine; UW 
116076-116090, 14 (63.6-79.8 mm), Puget Sound, 
48.7136°N, 123.0161°W, 60-80 m depth, RV Centennial, 
site 31, sample V21, 6 November 2006, D. Gunderson 
and J. Blaine; UW 118502, 23 (68-83 mm), Salish Sea, 
San Juan Channel, south of Turn Island, 48.52°N, 
122.95°W, 73-77 m depth, 24 April 2007, L. L. Britt; 
UW 118503, 8 (68.0-85.0 mm), Salish Sea, San Juan 
Channel, Jackson Beach, 48.52°N, 123°W, beach seine, 
24 April 2007, L. L. Britt; UW 16754, 1 (125 mm), 
Puget Sound, Golden Gardens, 47.7°N, 122. 4°W, 1 Jan- 
uary 1963, R. B. Grinols; UW 26833, 11 (102.6-121.6 
mm), Salish Sea, San Juan Island, Eagle Cove, 48.5°N, 
123°W, Friday Harbor Lab 75-03-02, 20 June 1975, 
Fish 454a class. California: HSU 4802, 34 (77-108 
mm), Humboldt Bay, 40.7621°N, 124.2152°W, 1 October 
2009, A. Kinziger; UW 152676, 104 (60-77 mm), Hum- 
Appendix 2 
This appendix provides diagnostic mitochondrial DNA 
sequence data for North Pacific sand lances of the 
genus Ammodytes, including data for neotypes of A. 
hexapterus and A. personatus and for the holotype of 
A. heinan new species. The sequences are a partial 
boldt Bay, north jetty, 40.7621°N, 124.2152°W, 1 Octo- 
ber 2009, A. Kinziger. 
Ammodytes japonicus 138 specimens (30.0-160.9 
mm) examined, including the types listed in the spe- 
cies description in the main text. FAKU 130765, 1 
(104.8 mm), Japan, Nohara, Maizuru, Kyoto, 35.5°N, 
135. 4°E; FAKU 130979-131008, 30 (131.2-160.9 mm), 
Japan, Inland Sea, off Akashi, Hyogo, 34.5°N, 135°E, 
Y. Kai; FAKU 131055-131091, 28 (86.0-108.8 mm), 
Japan, off Shiroko, Aichi, Ise Bay, 34.734°N, 136.68°E, 
Y. Kai; FAKU W856, W914, W915, 125465, 130765, 
5 (103.7-167.0 mm), Japan, Sea of Japan, Kyoto, 
Wakasa Bay, 35.8°N, 135.5°E, 1977; UW 153060, 17 
(30.0-45.0 mm), Japan, Maizuru, 35.5°N, 135. 4°E; 
UW 153061, 31 (30-32 mm), Japan, Ise Bay, 34.7°N, 
136. 8°E, 2006; UW 152684, 3 (208.0-218.9 mm), Ja- 
pan, off Wakkanai, Hokkaido, 45.4492°N, 141.6446°E, 
2007, Y. Kai; UW 16457, 5 (118.5-146.8 mm), Japan, 
Shizuoka Province, Suruga Bay, 35°N, 138.75°E, 15 
May 1950, F. Wilke. 
Ammodytes dubius 11 specimens (87.0-125.0 mm) ex- 
amined. UW 152011, 10 (87-125 mm), western North 
Atlantic Ocean, Massachusetts, 41.7637°N, 69.8956°W; 
UW 4799, 1 (115 mm), Woods Hole, Massachusetts, 2 
October 2012. 
Ammodytes americanus 3 specimens (73.0-142.0 mm) 
examined. VIMS 12840, 1 (142 mm), western North 
Atlantic Ocean, Rhode Island Sound, 41.2355°N, 
71.5519°W, 29 m depth, FV Darana R, cruise 
NM20100401, station 150, 6 May 2010; VIMS 12843, 
1 (138 mm), western North Atlantic Ocean, Rhode Is- 
land Sound, 41.1764°N, 71.4936°W, 29 m depth, FV 
Darana R, cruise NM20100401, station 139, 5 May 
2010; VIMS 12844, 1 (73 mm), western North Atlantic 
Ocean, Rhode Island Sound, 41.2937°N, 71.8408°W, 31 
m depth, FV Darana R, cruise NM20 100401, station 
126, 11 May 2010. 
Ammodytes marinus 2 specimens (133.0-155.0 mm) 
examined. ZMUC P61825-P61826, 2 (133-155 mm), 
Denmark, North Sea, Horns Rev, 55.47°N, 8.43°E, 2 
July 2009. 
Ammodytes tobianus 2 specimens (105.0-106.0 mm) 
examined. ZMUC P61815-P61816, 2 (105-106 mm), 
Denmark, Langeland, off Hjortholm, 55.9°N, 10.6°E, 1 
May 2010. 
(560-base-pair) fragment from the cytochrome oxidase 
c subunit 1 region of the mitochondria. 
Cytochrome oxidase c subunit 1 sequences 
Ammodytes hexapterus 
UAM 2813, neotype, GenBank Accession KJ137280. 
