Fishery Bulletin 96(4), 1998 
provides a simplified yet phylogenetically informative 
framework for future studies involving this family. 
Taxonomic list of the PSeuroneetidae 
Subfamily Hippoglossinae 
Genus Reinhardtius (R. evermanni, 2 
R. hippoglossoides, R. stomias ) 
Genus Hippoglossus ( H . hippoglossus, H. stenolepis) 
Genus Verasper (V. moseri, V. variegatus ) 
Genus Clidoderma ( C . aspert'imum 2 ) 
Subfamily Eopsettinae 
Genus Eopsetta (E. grigorjewi, E.jordani ) 
Subfamily Lyopsettinae 
Genus Lyopsetta (L. exdis) 
Subfamily Hippoglossoidinae 
Genus Acanthopsetta (A. nadeshnyi ) 
Genus Cleisthenes (C. herzensteini, C. pinetorum) 
Genus Hippoglossoides ( H . dubius, 2 H. elassodon, 
H. platessoides, H. robustus) 
Subfamily Pleuronectinae 
Tribe Psettichthyini 
Genus Psettiehthys (P. melanostictus) 
Tribe Isopsettini 
Genus Isopsetta (I. isolepis) 
Tribe Microstomini 
Genus Lepidopsetta ( L . bilineata, L. mochigarei) 
Genus Dexistes ( D . rikuzenius) 
Genus Pleuronichthys ( P . coenosus, 2 P. cornutus, 
P. decurrens, P. guttulatus, P. ocellatus, P. ritteri, 
P. uerticalis) 
Genus Microstornus (M. achne, M. bathybius, 
M. kitt, M. pacificus, M. shuntovi 2 ) 
Genus Glyptocephalus ( G . cynoglossus , G. kitaharai, 
G. stelleri , G. zachirus) 
Tribe Pleuronectini 
Genus Parophrys (P. vetula ) 
Genus Limanda 3 (L. aspera, L. ferruginea, 
L. limanda , L. proboscidea, L. punctatissima, 
L. sakhalinensis ) 
Genus Platiehthys (P. bicoloratus, P. flesus, 
P. stellatus ) 
Genus Pleuronectes (P. glacialis, P. pinnifasciatus, 
P. platessus, P . putnami, P. quadrituberculatus) 
Genus Pseudopleuronectes ( P . americanus, 
P. herzensteini, P. obscurus, 2 P. schrenki, 
P.yokohamae) 
2 Species not included in analysis, phylogenetic position and clas- 
sification determined a posteriori. 
3 Monophyletic status is uncertain. 
The reclassification of several species is necessary 
to incorporate the new phylogenetic information. The 
changes to the classification are as follows: 
1 Atherestes evermanni, A. stomias, and Rein- 
hardtius hippoglossoides are united within 
Reinhardtius which is the oldest valid genus- 
group name. Reinhardtius (=Atherestes) ever- 
manni was not examined and its position within 
the subfamily Hippoglossinae was determined a 
posteriori. Placement of R. evermanni within the 
clade uniting these three species is based on ex- 
amination of external morphological characters, 
number of abdominal vertebrae, and the descrip- 
tion provided in Norman (1934). 
2 The genus Eopsetta ( sensu Sakamoto, 1984a) with 
E. exilis, E. grigorjewi, and E. jordani is para- 
phyletic. The revised Eopsetta contains E. grigorjewi 
and E. jordani whereas E. exilis, is reassigned to 
the genus Lyopsetta as in Norman (1934). 
3 Hippoglossoides herzensteini and H. pinetorum 
( sensu Sakamoto, 1984a) are reclassified as 
Cleisthenes, as in Norman (1934). This is a sub- 
jective reclassification because interrelationships 
of these two species within Hippoglossoidinae 
does not contradict the previous classification. It 
is based on the monophyletic status of these two 
species and maintains a hierarchically consistent 
nomenclature within the family. 
4 A number of revisions were made to produce a 
simplified nomenclature that is hierarchically 
consistent with the phylogenetic interrelation- 
ships. Species of Hypsopsetta and Pleuronichthys 
( sensu Sakamoto, 1984a) are regrouped under 
Pleuronichthys. The species-group name (guttu - 
lata) is revised to agree with gender in the genus 
(i.e. Pleuronichthys guttulatus ). The species of 
Embassichthys and Microstornus ( sensu Saka- 
moto, 1984a) are regrouped under Microstornus. 
The species of Errex, Glyptocephalus, and Tanakius 
( sensu Sakamoto, 1984a) are regrouped under 
Glyptocephalus . The species of Kareius and Pla- 
tichthys ( sensu Sakamoto, 1984a) are regrouped 
under Platiehthys. 
5 The genus Pleuronectes ( sensu Sakamoto, 1984a), 
with 19 species, is not monophyletic. According 
to our results Pleuronectes comprises only five 
species: P. glacialis, P . pinnifasciatus, P. platessus, 
P. putnami, and P. quadrituberculatus . The spe- 
cies-group name (platessa ) is revised to corre- 
spond with gender in the genus (i.e. Pleuronectes 
platessus). Other species of Pleuronectes (sensu 
Sakamoto, 1984a) are reclassified as Lepidopsetta 
bilineata, L. mochigarei , Parophrys vetula, 
Limanda aspera, L. ferruginea, L. limanda, L. 
