858 
Fishery Bulletin 96(4), 1 998 
and European populations. Can. J. Fish. Aquat. Sci. 
52:1863-1872. 
McPeek, M. A., and R. D. Holt. 
1992. The evolution of dispersal in spatially and tempo- 
rally varying environments. Am. Nat. 140:1010-1027. 
McPhail, J. D. 
1997. The origin and speciation of Oncorhynchus revisited. 
In D. J. Stouder, P. A. Bisson and R. J. Naiman (eds.). Pa- 
cific salmon and their ecosystems: status and future op- 
tions, p. 29-38. Chapman and Hall, New York, NY. 
McPhail, J. D., and C. C. Lindsey. 
1970. Freshwater fishes of northwestern Canada and 
Alaska. Bull. Fish. Res. Board Can. 173, 381 p. 
1986. Zoogeography of freshwater fishes of Cascadia (the 
Columbia system and rivers north to the Stikine). In C. 
H. Hocutt and E. O. Wiley (eds.), Zoogeography of North 
American freshwater fishes, p. 615-637. John Wiley & 
Sons, Inc., New York, NY. 
Miller, K. M., and R. E. Withler. 
1997. Mhc diversity in Pacific salmon: population structure 
and trans-species allelism. Hereditas 127:83-95. 
Miller, K. M., R. E. Withler, and T. D. Beacham. 
1996. Stock identification of coho salmon ( Oncorhynchus 
kisutch ) using minisatellite DNA variation. Can. J. Fish. 
Aquat. Sci. 53:181-195. 
Nehlsen, W., Williams, J. E., and J. A. Lichatowich. 
1991 Pacific salmon at the crossroads: stocks at risk from 
California, Oregon, Idaho, and Washington. Fisheries 
16(2):4-21. 
Northcote, T. G., and D. Y. Atagi. 
1997. Pacific salmon abundance trends in the Fraser river 
watershed compared with other British Columbia systems. 
In D. J. Stouder, P. A. Bisson, and R. J. Naiman (eds.), Pacific 
salmon and their ecosystems: status and future options, p. 
199-219. Chapman and Hall, New York, NY. 
Pulliam, H. R. 
1988. Sources, sinks, and population regulation. Am. Nat. 
132:652-661. 
Pulliam, H. R., and B. J. Danielson. 
1991. Sources, sinks, and habitat selection — a landscape per- 
spective on population dynamics. Am. Nat. 137:S50-S66. 
Raymond, M., and F. Rousset 
1995a. GENEPOP (version 1.2): population genetics software 
for exact tests and ecumenism. Hereditary 86:248-249. 
1995b. An exact test for stock differentiation. Evolution 
49:1280-1283. 
Reisenbichler, R. R. 
1997. Genetic factors contributing to declines of anadro- 
mous salmonids in the Pacific northwest. In D. J. Stouder, 
P. A. Bisson, and R. J. Naiman (eds.), Pacific salmon and 
their ecosystems: status and future options, p. 223- 
224. Chapman and Hall, New York, NY. 
Ricker, W. E. 
1972. Heredity and environmental factors affecting certain 
salmonid populations. In R. C. Simon and P. A. Larkin (eds.), 
The stock concept of Pacific salmon, p. 19-60. H. R. MacMillan 
lectures in fisheries, Univ. British Columbia, Vancouver. 
Roff, D. A., and P. Bentzen. 
1989. The statistical analysis of mitochondrial DNA poly- 
morphisms: x 2 and the problem of small sample sizes. 
Mol. Biol. Evol. 6:539-545. 
Saitou, N., and M. Nei. 
1987. The neighbor-joining method: a new method for recon- 
structing phylogenetic trees. Mol. Biol. Evol. 4:406-425. 
Sakai, R. K., S. Scharf, F. Falloona, K. B. Mullis, 
G. T. Horn, H. A. Erlich, and N. Arnheim. 
1985. Enzymatic amplification of Beta-globin genomic se- 
quences and restriction site analysis for diagnosis of sickle 
cell anemia. Science (Wash., D.C.) 230:1350-1354. 
SAS Institute Inc. 
1989. SAS/STAT user’s guide, version 6, 4th edition, vol. 1. 
p. 678-771. SAS Institute Inc., Cary, NC. 
Scribner, K. T., J. R. Gust, and R. L. Fields. 
1996. Isolation and characterization of novel salmon 
microsatellite loci: cross-species amplification and popu- 
lation genetic applications. Can. J. Fish. Aquat. Sci. 
53:833-841. 
Small, M. P., T. D. Beacham, R. E. Withler, and 
R. J. Nelson. 
1998. Discriminating coho salmon (Oncorhynchus kisutch ) 
populations within the Fraser River, British Columbia. Mol. 
Ecol. 7:141-155. 
Tautz, D. 
1989. Hypervariability of simple sequences as a general 
source for polymorphic DNA markers. Nuc. Acid Res. 
17:6563-6471. 
Van Doornik, D. M., G. B. Milner, and G. A. Winans. 
1996. Transferrin polymorphism in coho salmon, Oncorhy- 
nchus kisutch , and its application to genetic stock 
identification. Fish. Bull. 94:566-575. 
Walters, C. J. 
1993. Where have all the coho gone? In L. Berg and P. W. 
Delaney (eds.), Proceedings of the coho salmon workshop, 
Nanaimo, B.C., May 26-28, 1992, p. 1-8. Department of 
Fisheries and Oceans, Nanaimo. 
Waples, R. S. 
1991. Pacific salmon, Oncorhynchus spp., and the defini- 
tion of “species” under the Endangered Species Act. Mar. 
Fish. Rev. 53(3):ll-22. 
Warner, B. G., R. W. Mathewes, and J. J. Clague. 
1982. Ice-free conditions on the Queen Charlotte Islands, 
British Columbia, at the height of late Wisconsin glacia- 
tion. Science (Wash. D.C.) 218:675-677. 
Wehrhahn, C. F., and R. Powell. 
1987. Electrophoretic variation, regional differences and 
gene flow in the coho salmon ( Oncorhynchus kisutch) of 
Southern British Columbia. Can. J. Fish. Aquat. Sci. 
44:822-831. 
Weir, B. S., and C. C. Cockerham. 
1984. Estimating F-statistics for the analysis of population 
structure. Evolution 38:1358-1370. 
Weitkamp, L. A., T. C. Wainwright, G. J. Bryant, G. B. 
Milner, D. J. Teel, R. G. Kope, and R. S. Waples. 
1995. Status review of coho salmon from Washington, Or- 
egon, and California. U.S. Dep. Commerce, NOAA Tech. 
Memo. NMFS-NWFSC-24, 258 p. 
Wood, C. C., B. E. Riddell, D. T. Rutherford, and 
R. E. Withler. 
1994. Biochemical genetic survey of sockeye salmon 
(Oncorhynchus nerka) in Canada. Can. J. Fish. Aquat. 
Sci. 51:114-131. 
