862 
Fishery Bulletin 96(4), 1998 
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Figure 1 
Micrographs of testicular developmental stages in male English sole: (A) regressed (510x), (B) early recrudescence (560x), 
(C) late recrudescence (560x), (D) early spermiogenesis (560x), (E) spawning (540x), and (F) spawned out ( 180x). a=spermatogonia, 
b=primary spermatocytes, c=secondary spermatocytes, d=spermatids, and e=spermatozoa. 
enter spermiogenesis. However, only a small propor- 
tion of males <230 mm were actually producing 
sperm, and none in this size range were collected on 
the spawning ground. The smallest male caught at 
University Point measured 238 mm, suggesting that 
males <230 mm may not migrate to the spawning 
ground (University Point) (Table 3). 
Changes in reproductive parameters 
during gonadal recrudescence 
Female Figure 4, A and B, shows reproductive pa- 
rameters in female sole at each developmental stage. 
Generally, the immature (regressed and previ- 
tellogenic) sole had low GSI (1.4 ±0.1, 85), and low 
