U.S. Department 
of Commerce 
Carlos M. Gutierrez 
Secretary 
National Oceanic 
and Atmospheric 
Administration 
Vice Admiral 
Conrad C. Lautenbacher Jr., 
USN (ret.) 
Under Secretary for 
Oceans and Atmosphere 
National Marine 
Fisheries Service 
John Oliver 
Acting Assistant Administrator 
for Fisheries 
Scientific Editor 
Adam Moles, Ph.D. 
Associate Editor 
Elizabeth Siddon 
Ted Stevens Marine Research Institute 
Auke Bay Laboratories 
Alaska Fisheries Science Center 
17109 Pt. Lena Loop Road 
Juneau, Alaska 99801 
Managing Editor 
Sharyn Matriotti 
National Marine Fisheries Service 
Scientific Publications Office 
7600 Sand Point Way NE, BIN C15700 
Seattle, Washington 98115-0070 
The Fishery Bulletin (ISSN 0090-0656) 
is published quarterly by the Scientific 
Publications Office, National Marine 
Fisheries Service, NOAA, 7600 Sand 
Point Way NE, BIN C15700, Seattle, WA 
98115-0070. Periodicals postage is paid 
at Seattle, WA. POSTMASTER: Send 
address changes for subscriptions to Fish- 
ery Bulletin, Superintendent of Docu- 
ments, Attn.: Chief, Mail List Branch, 
Mail Stop SSOM, Washington, DC 20402- 
9373. 
Although the contents of this publica- 
tion have not been copyrighted and may 
be reprinted entirely, reference to source 
is appreciated. 
The Secretary of Commerce has deter- 
mined that the publication of this peri- 
odical is necessary according to law for 
the transaction of public business of this 
Department. Use of funds for printing of 
this periodical has been approved by the 
Director of the Office of Management and 
Budget. 
For sale by the Superintendent of 
Documents, U.S. Government Printing 
Office, Washington, DC 20402. Subscrip- 
tion price per year: $36.00 domestic and 
$50.40 foreign. Cost per single issue: 
$21.00 domestic and $29.40 foreign. See 
back for order form. 
Editorial Committee 
Jeffrey M. Leis 
Thomas Shirley 
David Somerton 
Mark Terceiro 
Australian Museum, Sydney, Australia 
Texas A&M University 
National Marine Fisheries Service 
National Marine Fisheries Service 
Fishery Bulletin web site: www.fishbull.noaa.gov 
The Fishery Bulletin carries original research reports and technical notes on investigations in 
fishery science, engineering, and economics. It began as the Bulletin of the United States Fish 
Commission in 1881; it became the Bulletin of the Bureau of Fisheries in 1904 and the Fishery 
Bulletin of the Fish and Wildlife Service in 1941. Separates were issued as documents through 
volume 46; the last document was No. 1103. Beginning with volume 47 in 1931 and continuing 
through volume 62 in 1963, each separate appeared as a numbered bulletin. A new system 
began in 1963 with volume 63 in which papers are bound together in a single issue of the 
bulletin. Beginning with volume 70, number 1, January 1972, the Fishery Bulletin became a 
periodical, issued quarterly. In this form, it is available by subscription from the Superintendent 
of Documents, U.S. Government Printing Office, Washington, DC 20402. It is also available free in 
limited numbers to libraries, research institutions. State and Federal agencies, and in exchange 
for other scientific publications. 
