Orr and Hawkins: Species of the rougheye rockfish complex 
113 
Table 1 
Proportional morphometries and meristics of Sebastes aleutianus (rougheye rockfish) and Sebastes melanostictus (blackspotted 
rockfish) for specimens in which all characters were used for multivariate analyses. Morphometric data are given in percent SL 
(standard length) and presented as the range, followed by the mean ± standard deviation (SD). P values are reported for sta- 
tistically significant differences at a level of 0.05, as evaluated by ANCOVA or Kruskal-Wallis tests when appropriate; ns = not 
statistically significant at an a level of 0.05. 
Sebastes aleutianus 
Sebastes melanostictus 
Range (mean ±SD) 
Range (mean ±SD) 
P 
n 
57 
80 
Standard length (mm) 
77.1-470 
101.6-445 
Meristics 
Dorsal-fin rays 
12-14 (13.4 ±0.5) 
13-15 (13.8 ±0.5) 
<0.0001 
Anal-fin rays 
6-8(7 ±0.3) 
6-8(7 ±0.2) 
ns 
Pectoral-fin rays (left) 
17-19 (18.1 ±0.3) 
17-19 (18.1 ±0.3) 
ns 
Lateral-line pores (left) 
30-34 (31.2 ±0.9) 
30-36(32 ±1.2) 
<0.0001 
Gill rakers 
29-33 (31.1 ±1.0) 
30-35 (33 ±1.1) 
<0.0001 
Morphometries 
Head length 
34.6-42.5 (37 ±1.6) 
34.6-41.4 (37.6 ±1.5) 
Orbit length 
8.8-13.5 (10.7 ±1.1) 
8.2-13.3 (10.4 ±1.0) 
ns 
Snout length 
5. 8-9. 7 (7.5 ±0.7) 
6. 6-9. 5 (7.9 ±0.6) 
0.0486 
Interorbital width 
5.5-8. 9 (7.1 ±0.7) 
5. 6-8. 8 (7.4 ±0.7) 
Suborbital depth 
1.6-2. 9 (2.3 ±0.3) 
1.5-3. 3 (2.3 ±0.4) 
0.0019 
Upper jaw length 
16.9-21.9 (19.2 ±0.9) 
17.2-21.5 (19.2 ±0.7) 
Lower jaw length 
20.4-27.2(23.4 ±1.2) 
20.7-26.8(24.5 ±1.1) 
Gill raker length 
3. 4-6.1 (4.8 ±0.6) 
3. 9-6. 8 (5.6 ±0.5) 
<0.0001 
Depth at pelvic-fin base 
31.6-42.6(36 ±1.9) 
31.3-39.7 (34.9 ±1.8) 
Depth at anal-fin origin 
23.9-31.9(28 ±1.6) 
23.5-29.4 (26.2 ±1.5) 
<0.0001 
Depth at anal-fin insertion 
11.9-15.3 (13.9 ±0.7) 
11.3-15.3 (13.3 ±1.0) 
<0.0001 
Dorsal-fin spine I length 
4. 6-7. 2 (5.8 ±0.6) 
64-9.6(7.9 ±0.8) 
<0.0001 
Dorsal-fin spine IV length 
10.6-15 (13.2 ±0.9) 
12.3-18.9 (14.8 ±1.3) 
<0.0001 
Pectoral-fin base depth 
8.4-10.4(9.4 ±0.4) 
8.2-10.4 (9.5 ±0.4) 
ns 
Pectoral-fin ray length 
25-31 (27.4 ±1.3) 
22-30 (26.3 ±1.4) 
0.0013 
Pelvic-fin ray length 
19.3-25.1 (22.3 ±1.1) 
19.4-25 (21.5 ±1.2) 
0.0298 
Pelvic-fin spine length 
11.1-15.5 (13.5 ±1.2) 
11.4-16.9 (13.8 ±1.2) 
<0.0001 
Anal-fin spine I length 
4. 7-9. 4 (6.6 ±1.0) 
4.7-10.1 (7.1 ±1.2) 
<0.0001 
Anal-fin spine II length 
11-18.4(14.4 ±1.8) 
11.8-20.9 (15.2 ±2.2) 
<0.0001 
Anal-fin spine III length 
14.2-19.8 (16.5 ±1.3) 
12.4-20.9 (16.6 ±1.7) 
<0.0001 
Caudal peduncle depth 
7.8-10.7 (9.7 ±0.5) 
8.2-10.6 (9.2 ±0.5) 
Caudal peduncle dorsal length 
10.5-15.3 (12.2 ±0.9) 
11.3-15 (13 ±0.8) 
Caudal peduncle ventral length 
17.8-21.9 (19.6 ±0.9) 
17.4-21.7 (20.1 ±0.9) 
Preanal length 
67.1-77.6 (71.8 ±2.5) 
63.2-78.5 (70 ±2.6) 
<0.0001 
Predorsal length 
31.2-39.5 (34.3 ±1.9) 
31.6-38.4 (35 ±1.4) 
Spinous dorsal-fin-base length 
31.9-41.7 (35.5 ±2.3) 
32.4-40.9 (36.3 ±2.0) 
ns 
Soft dorsal-fin-base length 
20.7-25.6 (22.9 ±1.2) 
17.7-24.7 (21.5 ±1.4) 
<0.0001 
Anal-fin-base length 
13.3-18.2 (15.3 ±0.9) 
12.4-17 (14.2 ±1.0) 
ns 
Pre-pelvic-fin length 
37.8-53.1 (44.4 ±4.2) 
36.6-50.5 (42.4 ±2.5) 
morphometric characters were also tested for sexual 
dimorphism, where gender was treated as a categorical 
factor and the gender-species interaction was tested. 
Arcsine-transformed morphometric ratios (with SL or 
head length [HL] as denominator) were tested to meet 
the assumptions of normality and equality of variance 
required for ANCOVA. The following characters exhib- 
ited normal distributions and did not differ significantly 
in variance between species and thus were subjected to 
ANCOVA: orbit length, snout length, suborbital depth, 
gill-raker length, body depth at anal-fin origin, body 
depth at anal-fin insertion, dorsal-fin spine 1 length, 
dorsal-fin spine 4 length, pectoral-fin base depth, pec- 
toral-fin ray length, pelvic-fin ray length, pelvic-fin 
