Orr and Hawkins: Species of the rougheye rockfish complex 
123 
Table 5 (continued) 
Left infraorbital 2 spines 
0 
1 
2 
3 
4 
5 
6 
n 
Sebastes aleutianus 
Lower West Coast 
11 
15 
4 
1 
Gulf of Alaska 
1 
14 
25 
10 
2 
3 
Bering Sea and Aleutian Is. 
1 
9 
2 
1 
Total 
1 
26 
49 
16 
3 
4 
99 
Sebastes melanostictus 
Lower West Coast 
1 
5 
6 
2 
1 
1 
Gulf of Alaska 
4 
14 
31 
13 
9 
3 
1 
Bering Sea and Aleutian Is. 
3 
16 
20 
4 
1 
1 
Western Pacific Ocean 
1 
7 
2 
1 
Total 
8 
36 
64 
21 
12 
3 
3 
147 
Left infraorbital 3 spines 
0 
1 
2 
3 
4 
5 
6 
7 
n 
Sebastes aleutianus 
Lower West Coast 
18 
12 
1 
Gulf of Alaska 
26 
25 
4 
Bering Sea and Aleutian Is. 
2 
10 
1 
Total 
46 
47 
6 
99 
Sebastes melanostictus 
Lower West Coast 
7 
8 
1 
Gulf of Alaska 
3 
27 
38 
6 
1 
Bering Sea and Aleutian Is. 
1 
18 
22 
2 
1 
1 
Western Pacific Ocean 
3 
7 
1 
Total 
4 
55 
75 
10 
2 
0 
0 
1 
147 
orbit length; Fig. 5A) to spine IV (9.8-15.5% SL; Fig. 
5B; tips of dorsal-fin spines broken in lectotype) and 
decreasing in height to spine XII; spine XIII much lon- 
ger, forming anterior support of dorsal-fin rays; mem- 
branes of spinous dorsal fin moderately incised, less so 
posteriorly; soft dorsal fin with anterior rays longest, 
posterior rays gradually shortening. Anal-fin spine II 
shorter than or equal to III (11.0-18.4 vs. 13.0-20.2% 
SL; tip of anal-fin spine II broken in lectotype), rela- 
tively longer in juveniles, soft rayed portion of anal fin 
with anterior rays longest, posterior rays gradually 
shortening, posterior margin perpendicular to body axis 
or with slight posterior slant, anterior ray tips directly 
ventral to or forward of posterior tips, anterior tip of 
anal fin slightly rounded. Pectoral fins with ray 10 or 
11 longest, extending to or slightly anterior to vent, 
fin-ray length 24.4-31.0 (25.5) % SL; fin-base width 
8.4-10.4 (9.1) % SL; dorsalmost ray simple, ventral 
5-10 simple, others branched in adults; all rays simple 
in three juveniles less than 90 mm (63.4-77.1 mm), the 
smallest examined. Pelvic fins extending about 50-90% 
of distance from pelvic-fin base to anal-fin origin, fall- 
ing well short of vent, ray length 19.3-25.1 (22.8) % 
SL, spine length 51.7-76.0 (59.2) % ray length. All fin 
Table 6 
Factor loadings for principal component (PC) analysis 
of meristic characters of Sebastes aleutianus (rougheye 
rockfish) and Sebastes melanostictus (blackspotted rock- 
fish) in which all characters were used for multivariate 
analyses. 
PCI 
PC2 
PC3 
Dorsal-fin rays 
0.5620 
0.2917 
-0.0630 
Anal-fin rays 
0.2956 
0.5901 
-0.5291 
Pectoral-fin rays (left) 
0.1390 
-0.6710 
-0.5828 
Lateral-line pores (left) 
0.4905 
-0.1233 
0.6118 
Gill rakers 
0.5804 
-0.3181 
-0.0470 
spines exhibit allometric growth, and juveniles have 
relatively longer spines. Caudal fin shallowly emargin- 
ate, length 20.0-28.1% SL (fin damaged in lectotype). 
Vent positioned below dorsal-fin spine 10-11, 7-21 (9.4) 
% HL from anal-fin origin. 
Lateral body scales often with many (ca. 5-7) acces- 
sory scales in posterior field. Maxilla and underside of 
