Orr and Hawkins: Species of the rougheye rockfish complex 
125 
ered part of the geographic area now generally accepted 
to extend eastward only to Unimak Pass. 
Remarks 
The four syntypes of Sebastodes aleutianus were depos- 
ited at the USNM and CAS. The single USNM specimen 
(USNM 48800) was labeled “cotype” and is hereby desig- 
nated as the lectotype of Sebastes aleutianus. Among the 
syntypes, this was the only specimen illustrated (Jordan 
and Gilbert, 1899, plate 48). The three specimens at CAS 
(SU 12928) are designated paralectotypes. 
Individuals of S. aleutianus were apparently confused 
by Pallas (1814) as the “summer form” of S. variabilis 
(Pallas, 1814) (Jordan and Evermann, 1898), the dusky 
rockfish, one of the first species of Sebastes described 
from the Pacific Ocean — the first being its sister-species 
S. ciliatus (Tilesius, 1813), the dark rockfish (Kend- 
all, 2000; Orr and Blackburn, 2004; Hyde and Vetter, 
2007). Only a single specimen (MNHN 8670) is extant 
from the material Pallas examined for his description 
of Perea variabilis, and it has been designated the lec- 
totype of S. variabilis (Orr and Blackburn, 2004). The 
only other specimens identified as Perea variabilis by 
Pallas were housed at ZMB and are missing, presumed 
to have been destroyed during the Second World War 
(Paepke and Fricke, 1992). 
Evermann and Goldsborough (1907) described Se- 
bastodes swifti from two juveniles collected in a bay 
Depth (m) 
Figure 8 
Depth (m) distribution of Sebastes aleutianus (rough- 
eye rockfish, open bar) and Sebastes melanostictus 
(blackspotted rockfish, closed bar) based on material 
examined. Each bar represents numbers of catalogued 
museum lots, which may contain one or many specimens. 
Sebastes melanostictus is found at deeper depths than 
S. aleutianus (ANOVA, F=14.98, df=l, P=0.002). 
of southeastern Alaska. Although the authors com- 
pared them to S. crameri (Jordan in Gilbert, 1897), the 
darkblotched rockfish, both specimens exhibited the 
diagnostic morphological characters and pale, unspot- 
160°E 170°E 180° 170°W 160°W 150°W 140°W 130°W 120°W 
Distribution of the rougheye rockfish complex (open circle) in the eastern Pacific Ocean 
and Bering Sea based on National Marine Fisheries Service, Alaska Fisheries Science 
Center, survey data for the years 1961 to 2005. The rougheye rockfish complex includes 
Sebastes aleutianus and S. melanostictus, rougheye and blackspotted rockfish, which 
were recorded in 7758 tows. Although catch records from the northwestern Bering Sea 
indicated an absence of both species, species of Sebastes other than S. alutus (Pacific 
ocean perch) were routinely identified only to genus. Most symbols represent more than 
one tow and capture. 
