Rhodes and Tupper: The vulnerability of Plectropomus areolatus to fishing 
199 
Table 2 
Temporal distribution of recaptured squaretail coralgrouper ( Plectropomus areolatus) taken outside (by the fishery) and inside 
(by researchers) the Kehpara Marine Sanctuary (KMS). The entire shaded area represents the reproductive season and the sales 
ban period is darkly shaded. Months are abbreviated. CP, total = cumulative percent of total catch by month; CP, ban = cumula- 
tive percent contribution of the total recaptures within the sales ban period, including those by fishermen and researchers. All = 
the combined recaptures by the fishery (outside the KMS) and by researchers (within the KMS); n = sample size of recaptures 
within the study period; CP sample size in) is that used to calculate cumulative percents; a = Peleng Channel or proximity; b = 
Dawak Channel or proximity; c = Nalap or vicinity; d = unknown locale; e = Liap; f = Penieu; g = Temmen. 
Year 
2005 
2006 
n 
Months 
Months 
J 
F 
M 
A 
M 
J 
J 
A 
S 
O 
N 
D 
J 
F 
Fishery 
Recaptures 
l a 
5 a.d 
7 a, C 
3 a,b 
ga,b,c 
-q 
CO 
0 
0 
l e 
l a 
6 a - d ' f 
0 
2 a 'g 
2 a 
39 
CP, total 
3 
15 
33 
41 
62 
69 
69 
69 
72 
74 
90 
90 
95 
100 
39 
CP, ban 
— 
— 
18 
26 
— 
10 
KMS 
Recaptures 
0 
2 
6 
5 
7 
20 
CP, total 
0 
10 
40 
65 
100 
20 
CP, ban 
— 
— 
30 
55 
— 
11 
All 
CP, total 
2 
14 
34 
49 
75 
80 
80 
80 
81 
83 
93 
93 
97 
100 
59 
CP, ban 
— 
— 
22 
36 
— 
21 
Table 3 
Presence (•) of acoustically tagged squaretail coralgrouper ( Plectropomus areolatus) to show the observed variability in resi- 
dency times (as site visitations) at the Kehpara Marine Sanctuary fish spawning aggregation (FSA) site over a 17-month period 
(January 2005-May 2006). The study period encompasses two complete reproductive seasons (~January-May). The table repre- 
sents only individuals present at the FSA for one month or more after initial tagging. 
J F 
M 
A 
M 
J 
J 
A 
S 
0 
N 
D 
J 
F 
M 
A M 
Males • • 
• 
• 
• 
• • 
• 
• • 
• 
• 
• 
• • 
• 
• • 
• 
• 
• 
• 
• 
• 
• 
• 
• 
• 
• 
• 
• • 
• 
• 
• • 
• 
• 
• 
• 
• 
• 
• 
• 
Females • 
• 
• 
• 
• 
• 
• 
• 
• • 
• 
• 
• 
• 
• 
• 
• 
• 
• 
Travel speed for three fish that moved between Peleng 
Channel and the KMS (=8 km) was 2.1 km/h (1 female, 
2 males), and all fish made the trip in <1 day. Although 
several acoustically tagged individuals moved south- 
ward to the KMS boundary, only one appeared to travel 
as far as Nalap. The apparent limited movement south- 
ward from the KMS is also reflected in the low number 
of conventional tag recaptures (n= 4) from areas south or 
east of KMS. No movement was detected directly within 
Kehpara Channel, in contrast to previous anecdotal re- 
ports of post-spawning migrations through the channel. 
Some individuals were detected seaward of the channel. 
