1953] 
Darlington — Australian Carabiclae 
59 
transversely sulcate in front, prosternal process very fine 
(attenuate) opposite front margins of coxae; prosternal 
episterna closely rugulose; indistinct traces of punctation 
at sides of prosternum (not on episterna) ; hind body below 
roughened at sides but not distinctly punctate. Inner wings 
reduced to narrow strips about V2 length of elytra; metepi- 
sterna shortened. Anterior femur with lower, posterior 
margin approximately straight (faintly convex) as seen 
from behind; anterior tibia externally with three long 
teeth and (above them) a small triangular projection; 
middle tibia with a slender spur 1/3 or i/4 from apex. Length 
5 mm. (or slightly less) ; width about 1.4 mm. 
Holotype (M. C. Z. No. 23,093) and 2 paratypes all from 
near the town of Margaret River, southwestern division 
of Western Australia, October, 1931. 
This species finds its closest relatives in the Heterogena 
Group of Clivina. In Sloane’s second key (1904, p. 714) it 
would go with olliffi SI. and blackhurni SI., differing from 
the former by being smaller and with narrower prothorax, 
and from the latter by having a distinct anterior transverse 
line on the pronotum. So far as I can judge from the 
descriptions alone, there are many other differences too. 
The separation of the median part of the clypeus from the 
clypeal wings in grata is so slight that the species might 
easily be referred to the Australasiae Group, where it would 
run (in Sloane’s key, 1904, p. 719) to ferruginea or to nigra 
or occulta, but it probably is not really very closely related 
to any of these. 
In my opinion, this new species {grata) really represents 
some stock close to heterogena Putz., modified as a result 
of reduction of the wings. The other characters which 
distinguish grata, especially the more oval elytra and 
shortened metepisterna, often accompany or follow wing- 
reduction among Carabidae {cf. Darlington 1936) and 
occur in some other (unhplated) flightless Australian 
Clivina, especially in the Procera Group. Even the reduced 
eyes of grata may be a secondary result of wing atrophy. 
I took, also at Margaret River, a single specimen of a 
Clivina which may represent the winged stock from which 
grata has been derived. It is bicolored like some examples 
